Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.
01526857
Claudio Contreras / Greater roadrunner (Geococcyx californianus) with nuptial gift calling mate, Laredo Borderlands, Texas, USA. April
00543021
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) at nest burrow, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
01636310
Hermann Brehm / Bee-eater (Merops apiaster) nuptial gift of insect prey, Hungary. June
01636307
Hermann Brehm / Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
01617044
Eric Medard / Sparrowhawk (Accipter nisus) pair mating, with dead chick, a nuptial gift, at female's feet, in forest, Pays de Loire, France
01617038
Eric Medard / Sparrowhawk (Accipter nisus) male flying to female. Dead chick, a nuptial gift, at female's feet, in forest, Pays de Loire, France
01617025
Eric Medard / Sparrowhawk (Accipter nisus) female with chick prey, nuptial gift from male, in forest, Pays de Loire, France
01617024
Eric Medard / Sparrowhawk (Accipter nisus) male and female with chick prey, nuptial gift from male, in forest, Pays de Loire, France
01617023
Eric Medard / Sparrowhawk (Accipter nisus) pair mating. With dead chick, a nuptial gift, at females feet, in forest, Pays de Loire, France
01613079
Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) on nest. Starfish (Asteroidea) nuptial gifts present around nest. Punta Albemarle, Isabela Island, Galapagos. June 2017.
01613078
Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) on nest. Starfish (Asteroidea) nuptial gifts present around nest. Punta Albemarle, Isabela Island, Galapagos. June 2017.
00583821
Sebastian Kennerknecht / Pheasant-tailed Jacana (Hydrophasianus chirurgus) flying in breeding plumage, Diyasaru Park, Colombo, Sri Lanka
01589948
Yashpal Rathore / Blue-tailed bee-eater (Merops philippinus) male offering female insect prey as nuptial gift, near Ranganathittu Bird Sanctuary, Karnataka, India.
01589947
Yashpal Rathore / Blue-tailed bee-eater (Merops philippinus) male offering female insect prey as nuptial gift, near Ranganathittu Bird Sanctuary, Karnataka, India.
01589882
Yashpal Rathore / Blue-tailed bee-eater (Merops philippinus) male bringing insect for female as nuptial offering. Near Ranganathittu Bird Sanctuary, Karnataka, India.
01589881
Yashpal Rathore / Blue-tailed bee-eater (Merops philippinus) male bringing insect for female as nuptial offering. Near Ranganathittu Bird Sanctuary, Karnataka, India.
01543518
Alex Hyde / Fishing spider (Trechalea) male with a silk-wrapped fish as a nuptial gift to offer to a female as part of his courtship behaviour, Central Caribbean foothills, Costa Rica.
01543517
Alex Hyde / Fishing spider (Trechalea) male with a silk-wrapped fish as a nuptial gift to offer to a female as part of his courtship behaviour, Central Caribbean foothills, Costa Rica.
01543516
Alex Hyde / Fishing spiders (Trechalea) male (left) offering female a silk-wrapped fish as a nuptial gift as part of his courtship behaviour, Central Caribbean foothills, Costa Rica
01543515
Alex Hyde / Fishing spiders (Trechalea) male (right) offering female a silk-wrapped fish as a nuptial gift as part of his courtship behaviour, Central Caribbean foothills, Costa Rica
01545805
Andres M. Dominguez / Kingfisher (Alcedo athis) male offering female small crayfish prey, Sierra de Grazalema Natural Park, southern Spain, April.
00543025
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543020
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) carrying fish prey, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
Bert Willaert / Close-up of the nuptial pads of a male Common toad (Bufo bufo) during the breeding season, photographed in controlled conditions, Belgium, March.
01117981
Christophe Courteau / West African giraffe male smelling female in oestrus {Giraffa camelopardis peralta}.
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Christophe Courteau / West African giraffe male smelling female in oestrus {Giraffa camelopardis peralta}.
00443833
Murray Cooper / Small Fruit Fly (Drosophilidae) group using mushroom as nuptial mating ground, Amazon, Ecuador
01062881
Jabruson / Shed termite wings {Isoptera} floating in puddle Tsavo NP, Kenya wings shed after nuptial flight
01029160
PREMAPHOTOS / Male Nursery web spider presents gift to female {Pisaura mirabilis} England. Many spiders have to bribe females into mating "nuptial gifts". She will terminate the contract when she's finished eating...
01106273
George McCarthy / Male Adders fighting in pre-nuptial display {Vipera berus} Surrey, UK
01106275
George McCarthy / Male Adders fighting in pre-nuptial display {Vipera berus} Surrey, UK
01106274
George McCarthy / Male Adders fighting in pre-nuptial display {Vipera berus} Surrey, UK
01106239
George McCarthy / Close up of foot showing nuptial pads of Common european toad {Bufo bufo} UK
00543034
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543033
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543032
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) group at coastal breeding colony, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543031
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) group at coastal breeding colony, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543028
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543027
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543026
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) carrying fish prey, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543024
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543023
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) at nest burrow at sunrise, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543022
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) in breeding plumage, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
00543019
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) carrying fish prey, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
21106272
George McCarthy / Male Adders fighting in pre-nuptial display {Vipera berus} Surrey UK
21100826
George McCarthy / Male Adders fighting in pre-nuptial fight {Vipera berus} Surrey UK
00543029
Sebastian Kennerknecht / Atlantic Puffin (Fratercula arctica) at sunrise, Skomer Island National Nature Reserve, Skomer Island, Pembrokeshire, Wales, United Kingdom
01196735
Jose Luis GOMEZ de FRANCISCO / Western marsh frog (Rana perezi), close-up of nuptial pad on foot of front leg, Spain
01719327
Mateusz Piesiak / White-winged tern (Chlidonias leucopterus) pair, courtship display, male presenting female with nuptial gift, Biebrza National Park, Poland. May.
01725207
David Pattyn / Green jay (Cyanocorax yncas) pair perched on branch, male giving food to female, a nuptial gift during courtship, Medellin area, Antioquia, Colombia.
01722223
Staffan Widstrand / Eurasian roller (Coracias garrulus) pair perched on branch, male delivering insect to female as a nuptial gift prior to mating, Kiskunsagi National Park, Pusztaszer, Hungary. May.
01702491
Joel Sartore / Photo Ark / Tricolor shiner (Cyprinella trichroistia) male with nuptial tubercles, Goodwater, Alabama. Captive.
01670297
Jussi Murtosaari / Arctic tern (Sterna paradisaea) male in flight with nuptial gift for female on rocks below. Hailuoto, Finland. May.
01653866
Doug Perrine / White tern (Gygis alba candida) pair, female with juvenile flying fish presented by mate as nuptial gift. Sand Island, Midway Atoll National Wildlife Refuge, Papahanaumokuakea Marine National Monument...
01653865
Doug Perrine / White tern (Gygis alba candida) pair, male presenting female with juvenile flying fish, a nuptial gift. Sand Island, Midway Atoll National Wildlife Refuge, Papahanaumokuakea Marine National Monument,...
01649195
John Chan / California least terns (Sternula antillarum browni) mating on the beach, with male holding nuptial gift of fish for female, Huntington Beach Least Tern Preserve, California, USA May
01649194
John Chan / California least terns (Sternula antillarum browni) mating on the beach, with male holding nuptial gift of fish for female, Huntington Beach Least Tern Preserve, California, USA May
01649193
John Chan / California least terns (Sternula antillarum browni) mating on the beach, male feeding female nuptial gift of fish, Huntington Beach Least Tern Preserve, California, USA May
01649192
John Chan / California least terns (Sternula antillarum browni) mating on the beach, female feeding on nuptial gift from male, Huntington Beach Least Tern Preserve, California, USA May
01649191
John Chan / California least terns (Sternula antillarum browni) mating on the beach - male feeding female nuptial gift. Huntington Beach Least Tern Preserve, California, USA May
01649190
John Chan / California least terns (Sternula antillarum browni) mating on the beach, with male holding nuptial gift of fish for female, Huntington Beach Least Tern Preserve, California, USA May
01649189
John Chan / California least terns (Sternula antillarum browni) mating on the beach, female feeding on nuptial gift from male, Huntington Beach Least Tern Preserve, California, USA May
01649188
John Chan / California least terns (Sternula antillarum browni) mating on the beach - male feeding female nuptial gift. Huntington Beach Least Tern Preserve, California, USA May
01649173
John Chan / California least terns (Sternula antillarum browni) mating on the beach - male feeding female nuptial gift. Huntington Beach Least Tern Preserve, California, USA May
01649172
John Chan / California least terns (Sternula antillarum browni) mating on the beach - male feeding female nuptial gift. Huntington Beach Least Tern Preserve, California, USA May