Will Burrard-Lucas / Tufted gray langur (Semnopithecus priam) Close up portrait of head and shoulders. Minneriya National Park, Sri Lanka. September

Ismaele Tortella / Eurasian otter (Lutra lutra) hunting in a shoal of Atlantic herring (Clupea harengus), northern Norway, March

Lucas Bustamante / Toque Macaques (Macaca sinica) Polonnaruwa, Central Province, Sri Lanka

Ernie Janes / Celebes crested macaque (Macaca nigra) infant, captive.

John Shaw / Snow monkey or Japanese macaque (Macaca fuscata) female with baby clinging to her back. Jigokudani, Japan, February.

Andy Rouse / Stonechat, (Saxicola rubicola) male shaking head, blurred motion. UK

SCOTLAND: The Big Picture / Otter (Lutra lutra) portrait on shore with boats in background. Argyll, Scotland, UK, August.

Bernard Castelein / Rhesus macaque (Macaca mulatta), mixed group of female and juveniles, walking along a track, Keoladeo National Park, Bharatpur, India

Bernard Castelein / Rhesus macaque (Macaca mulatta), juveniles playing in tree, Keoladeo NP, Bharatpur, India

Suzi Eszterhas / Cotton-top tamarin (Saguinus oedipus), adult with two week old baby on back. Northern Colombia.

Suzi Eszterhas / Red-shanked douc langur (Pygathrix nemaeus) infant sucking thumb while caretaker holds it. Animal was rescued from illegal wildlife trade, Endangered Primate Rescue Center, Cuc Phuong National Park, V...

David Pattyn / Bearded vulture (Gypaetus barbatus) in flight over winter landscape with snow, Leukerbad, Vallais, Wallis, Switzerland, January