John Waters / Female Grayling (Thymallus thymallus) pursued by several males, River Kennet, Berkshire, England, UK. March.

Michel Roggo / Grayling (Thymallus thymallus) on spawning ground. Lac de Fenetre, alpine lake, 2456 m above sea level, Canton of Valais, Switzerland, June.

Michel Roggo / Grayling (Thymallus thymallus) on spawning ground. Lac de Fenetre, alpine lake, 2456 m above sea level, Canton of Valais, Switzerland, June.

Michel Roggo / European Grayling (Thymallus thymallus), Male, Aare river, Switzerland, 2005

Michel Roggo / European Grayling (Thymallus thymallus) waiting for drift food, Sava river, Slovenia, 2004

Michel Roggo / European grayling (Thymallus thymallus) under waterfall waiting for drift food, Sava river, Slovenia, 2003

Michel Roggo / European Grayling (Thymallus thymallus) feeding, River Koridnica, Slovenia, 1999

Michel Roggo / European Grayling (Thymallus thymallus), Males at spawning ground, Lake of Thoune, Switzerland, 1992

Michel Roggo / European Graylings (Thymallus thymallus) spawning, male in foreground, Lake of Thoune, Switzerland, 1992

Michel Roggo / European Grayling (Thymallus thymallus), Under waterfall waiting for drift food, Baca river, Slovenia, 2006

Wild Wonders of Europe / Roggo / European grayling (Thymallus thymallus) pair spawning, the dark one is the male, Lake Thun, Thun, Switzerland, March 2009

Wild Wonders of Europe / Roggo / European grayling (Thymallus thymallus) Ain river, France, June 2009