Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

Rolf Nussbaumer / Dung Beetle (Scarabaeinae) adult rolling dung ball. Laredo, Webb County, South Texas, USA Not available for ringtone/wallpaper use.

Gavin Hellier / Tuk Tuk or auto rickshaw in motion at night, Bangkok, Thailand, 2010. Model released.

Kristel Richard / Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain.

Steven Kazlowski / Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA

Claudio Contreras / Heermann's Gull (Larus heermanni) in flight, Ensenada, Baja California Peninsula, Mexico, December.

Jordi Chias / Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011, Commended in Wildlife Photographer of the Yea...

Eric Medard / Bechstein's Bat (Myotis bechsteinii) in flight at night, mouth open to emit echolocating calls. France, Europe, September.

Carol Walker / Cowboy galloping through snow, wearing thick sheepskin coat, Wyoming, USA, February 2012, model released

Richard Du Toit / Springbok (Antidorcas marsupialis) silhouetted on the horizon. Namaqualand, South Africa, December.

Erlend Haarberg / Norway lemming (Lemmus lemmus) swimming, Sweden, May

Ernie Janes / Greylag geese (Anser anser) in flight with paraglider behind, Cley bank, Norfolk, UK

Andy Sands / Long Water Measurer (Hydrometra stagnorum) drinking by piercing surface tension with mouthparts. Captive, UK, July.

Eric Baccega / Muskrat (Ondatra zibethicus) walking on snow, Quebec, Canada

Gerrit Vyn / Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA

Nick Turner / Couple with child in seat cycling in the Cotswolds at Guiting Power, Gloucestershire, UK, August 2011. Model released

Paul Hobson / Feral Pigeon / Rock Dove (Columba livia) on city street. Sheffield, UK.

Alex Mustard / Yellowhead wrasse (Halichoeres garnoti) male performing a courtship dance to attract females, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Alex Mustard / Mirrorwing flying fish (Hirundichthys speculiger) mirrored and swimming beneath the surface at night, between Bahamas and Gulf Stream, West Atlantic Ocean, Florida, USA.

Carol Walker / Horse in snow storm with shed in background, USA

George Sanker / Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October

Edwin Giesbers / European mink (Mustela lutreola) swimming, captive, Germany, Critically endangered

Petra Wegner / Cavalier King Charles Spaniel, ruby, running out of the sea wearing a lifejacket / life vest.

Brent Stephenson / New Zealand Albatross (Diomedea antipodensis) in flight, showing upperwing, wingtip touching the sea surface. Off Kaikoura, Canterbury, New Zealand, October.

Brent Stephenson / Male Australasian Gannet (Morus serrator) flying with seaweed for use as nesting material. Cape Kidnappers, Hawkes Bay, New Zealand, September.

Todd Mintz / Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March

Todd Mintz / Narwhal (Monodon monoceros) diving sequence 1/4, Pond Inlet, Nunavut, Baffin Island, Canadian Arctic, June

Robin Chittenden / Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November

Philippe Clement / The harbour at Le Croisic, Loire-Atlantique, Pays-de-la-Loire, France September 2011

Markus Varesvuo / Spotted Redshank (Tringa erythropus) in flight, Utsjoki Finland June

Markus Varesvuo / Magpie (Pica pica) in flight, Sweden October

Pete Oxford / Caribbean trumpetfish (Aulostomus maculatus) Bonaire, Netherlands Antilles, Caribbean

Luis Quinta / Baskian viper (Vipera seoanei) Geres National Park, Portugal, September.

John Abbott / Eastern Green june beetle (Cotinis nitida) in flight, Texas, USA, October

John Abbott / Snakefly (Agulla sp) in flight, Wild Basin Preserve, Texas, USA, April

Loic Poidevin / Redshank (Tringa totanus) walking on mud. Breton Marsh, French Atlantic Coast, November.

Nick Upton / Great capricorn / Great oak longhorn beetle (Cerambyx cerdo), one of the largest beetles in Europe, on tree trunk, Lesbos (Lesvos), Greece, June.

Kristel Richard / A purebred Lusitano stallion galloping in the arena at the Companhia das Lezírias Stud Farm, Samora Correia, Santarém, Portugal, July 2011

Kristel Richard / A purebred Lusitano colt (with his mane and tail cut to mark his first year) trotting in the arena at the Companhia das Lezírias Stud Farm, Samora Correia, Santarém, Portugal, July 2011

John Abbott / Ichneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November

Nick Garbutt / Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November

Sven Zacek / House Martin (Delichon urbicum) in flight with invertebrates in its beak. Estonia, Europe, July.

Sven Zacek / Barn Swallow (Hirundo rustica) in flight. Estonia, Europe, June.

Sven Zacek / Ural Owl (Strix uralensis) in flight over snow. Virumaa, Estonia, Europe, February.

Sven Zacek / Roe Deer (Capreolus capreolus) pronking away from the photographer. Virumaa, Estonia, Europe, February.

Andy Rouse / Impala (Aepyceros melampus) male jumping during rut. Masai Mara, Kenya, Africa.

Matthew Maran / Chinook Salmon (Oncorhynchus tshawytscha) leaping through water surface in front of forested coastline. Vancouver Island, British Columbia, Canada, September.

Nick Garbutt / Acacia sp. at night, with a herd of Plains Zebra (Equus quagga) beneath. Near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania.

Robin Chittenden / Grass snake (Natrix natrix) swimming at surface, Dobrogea Tulcea, Danube Delta, Romania, May

Ole Jorgen Liodden / Brunnich's Guillemot (Uria lomvia) flying between misty colony cliffs. Spitsbergen, Svalbard, August.

Ole Jorgen Liodden / Black-legged Kittiwake (Rissa tridactyla) taking off from water. Spitsbergen, Svalbard, August.

Ole Jorgen Liodden / Oryx (Oryx gazella) in motion. Kalahari, Botswana, November.

Nuno Sa / Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August.

Nuno Sa / Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August.

Fabrice Cahez / Red fox (Vulpes vulpes) pouncing while hunting in field, Vosges, France, August

Fabrice Cahez / Female Roe deer (Capreolus capreolus) leaping, Vosges, France, April

Fabrice Cahez / Wild cat (Felis silvestris) pouncing in field, sequence 1/3, Vosges, France, May

Christophe Courteau / Whale headed / Shoebill stork (Balaeniceps rex) in flight over the swamps of Mabamba, Lake Victoria, Uganda / Bec en sabot, Lac Victoria, Ouganda

Kristel Richard / A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal.

Doug Allan / Icefish (Chaenocephalus aceratus) Signy Island, South Orkney Islands, Antarctica, Freeze Frame book plate page 92

Doug Allan / Minke Whale (Balaenoptera acutorostra) at the surface in vast seascape wilderness. Antarctica, February. Freeze frame book plate page 135.

Ernie Janes / Yellowhammer (Emberiza citinella) male in flight, Norfolk, UK, May

Ernie Janes / Combine harvester harvesting field of Wheat, Bedfordshire, UK, August 2006

Jan Hamrsky / Mosquito (Culex pipiens) adult emerging through water surface from larva. Europe, August.

Andrew Parkinson / Mute swan portrait on water (Cygnus olor) Derbyshire, UK, April.

Andrew Parkinson / Barn owl (Tyto alba) adult in flight in evening light, Norfolk, UK

Graham Eaton / Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea, seen from above. Cayenne, French Guiana, July.

Graham Eaton / Leatherback Turtle (Dermochelys coriacea) hatchling crossing a beach to get to the sea. Cayenne, French Guiana, July.

Kim Taylor / Brown Long-eared bat (Plecotus auritus) drinking from a lily pond , Surrey, UK. Overall WINNER of the Audubon Society of Greater Denver Share the View competition 2011

Stefan Huwiler / Blue Tit (Parus caeruleus) adult in flight. Zug, Switzerland, Europe, February.

Pete Oxford / Galapagos penguin (Spheniscus mendiculus) diving underwater, Bartolome Island, Galapagos, endemic.

Jerry Monkman / Woman mountain biking on a trail in an abandoned granite quarry on Millstone Hill, Barre, Vermont, USA. autumn. Model released, October 2010

Loic Poidevin / Man walking over colourful mineral deposits at the Dallol hydrothermal site on the Karoum salt lake, Danakil depression, northern Ethiopia, February 2009

Klaus Echle / Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November.

Klaus Echle / Red Fox (Vulpes vulpes) with wet fur in warm light. Black Forest, Germany, November.

Ernie Janes / House spider (Tegenaria domestica) on soap dish, UK

Frederique Olivier / Young girl paddling dugout canoe, Peava village, Solomon Islands, Melanesia, September 2008.

Rolf Nussbaumer / Cane Toad / Marine Toad / Giant Toad (Bufo marinus) adult jumping in Dogweed (Dyssodia pentachaeta) field. Laredo, Webb County, South Texas, USA, April.

Jeff Rotman / Poisonous Bluespotted ribbontail stingray (Taeniura lymma) at night, Perth, Australia, Indian Ocean.

Andy Rouse / Barn owl (Tyto alba) in flight over long grass, hunting, Wiltshire, UK

Andy Rouse / Short Eared Owl (Asio flammeus) hunting above grassland. UK, Europe, December.

Markus Varesvuo / Golden Eagle (Aquila chrysaetos) flying low over snow with talons brandished. Kuusamo, Finland, February.

Klaus Echle / After being trapped in an empty shed, Badger (Meles meles) passes wall decorated with graffiti on its way back to the forest. Baden-Wuerttemburg, Germany, May 2010. Winner of the Man and Nature catego...

Bence Mate / Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition.

Markus Varesvuo / Herring Gull (Larus argentatus) flying in snow Norway, March.

Solvin Zankl / Common european toad {Bufo bufo} two males attempting to mate with one female, Germany, March. Highly Commended, Other Animals category, GDT competition 2011

Klaus Echle / Badger (Meles meles) foraging in woodland on edge of forest, Black Forest, The Black Forest, Germany, May. Highly commended in Mammals category, GDT competition 2011

Klaus Echle / Red fox (Vulpes vulpes) vixen 'Sophie' became accustomed to the the photographer and would allow him to walk through forest beside her, Black Forest, Germany, July. Winner of the Fritz Polking portfol...

Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) in flight, Svalbard, Norway, July. Highly commended, BIRDS category, GDT 2011 competition. Joint runner up in Creative Visions of Nature Category, Wildlife Photographer...

Nuno Sa / Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins. Azores, North Atlantic, Septemb...

Anup Shah / Bonobo (Pan paniscus) adolescent male leaping through the trees, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo. October.

Fiona Rogers / Bonobo (Pan paniscus) adolescent male wading through water, calling, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo. October.

Ernie Janes / Barn owl (Tyto alba) in flight, viewed through car windscreen at night, Norfolk, UK, model released. Digital composite

David Woodfall / Red Kite (Milvus milvus) in flight. Gilfach farm SSSI, Radnorshire Wildlife Trust, Wales, January.

Fabrice Cahez / Red Fox (Vulpes vulpes) hunting in a frosty field. Vosges, France, October.

Fabrice Cahez / Muskrat (Ondatra zibethicus) at water surface. River Allier, France, April.