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Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan.
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Yukihiro Fukuda / Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan.
Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May
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Ross Hoddinott / Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May
Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m.
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David Shale / Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m.
Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July
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Eric Baccega / Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil. (This image may be licensed either as rights managed or royalty free.)
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Staffan Widstrand / RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil. (This image may be licensed either as rights managed or royalty free.)
Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.
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Morley Read / Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.
Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 2 of 3
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Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 2 of 3
Polar bear (Ursus maritimus) male walking across frozen landscape. Svalbard, Norway, April 2019.
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Danny Green / Polar bear (Ursus maritimus) male walking across frozen landscape. Svalbard, Norway, April 2019.
Polar bear (Ursus maritimus) male in snow. Svalbard, Norway, April.
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Danny Green / Polar bear (Ursus maritimus) male in snow. Svalbard, Norway, April.
RF - Polar bear (Ursus maritimus) male walking through snow covered landscape, mountains in background. Svalbard, Norway, April (This image may be licensed either as rights managed or royalty free.)
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Danny Green / RF - Polar bear (Ursus maritimus) male walking through snow covered landscape, mountains in background. Svalbard, Norway, April (This image may be licensed either as rights managed or royalty free.)
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December.
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Nick Garbutt / Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December.
Hawfinch (Coccothraustes coccothraustes) perched on branch amongst Hawthorn (Crataegus sp) berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February.
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Andres M. Dominguez / Hawfinch (Coccothraustes coccothraustes) perched on branch amongst Hawthorn (Crataegus sp) berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February.
American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba.
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Shane Gross / American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba.
Leopard seal (Hydrurga leptonyx) with mouth open, Antarctic Peninsula, Antarctica.
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Jordi Chias / Leopard seal (Hydrurga leptonyx) with mouth open, Antarctic Peninsula, Antarctica.
View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.
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Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.
Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.
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Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.
Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.
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Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.
Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive.
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Steven David Miller / Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive.
Aerial view of Fin whale (Balaenoptera physalus) lunge-feeding, with mouth open and throat pouch distended, southern Sea of Cortez (Gulf of California), Baja California, Mexico.
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Mark Carwardine / Aerial view of Fin whale (Balaenoptera physalus) lunge-feeding, with mouth open and throat pouch distended, southern Sea of Cortez (Gulf of California), Baja California, Mexico.
Giant panda (Ailuropoda melanoleuca) sitting with Bamboo in background. Shenshuping Panda Base, Wolong Nature Reserve, Wenchuan, Sichuan Province, China. Captive.
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Suzi Eszterhas / Giant panda (Ailuropoda melanoleuca) sitting with Bamboo in background. Shenshuping Panda Base, Wolong Nature Reserve, Wenchuan, Sichuan Province, China. Captive.
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.
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Franco Banfi / Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.
Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January.
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Eric Baccega / Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January.
Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August. Sequence 1/2.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August. Sequence 1/2.
Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August.
Emperor penguin (Aptenodytes forsteri) chick begging for food, on parent's feet. Atka Bay, Antarctica. August.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick begging for food, on parent's feet. Atka Bay, Antarctica. August.
Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
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Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
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Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
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Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.
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Paul Hobson / Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling, Montana
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Donald M. Jones / Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling, Montana
Western Meadowlark (Sturnella neglecta) male calling, Montana
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Donald M. Jones / Western Meadowlark (Sturnella neglecta) male calling, Montana
Elk (Cervus elaphus) bull bugling, Alberta, Canada
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Donald M. Jones / Elk (Cervus elaphus) bull bugling, Alberta, Canada
Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
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Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
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Paul Souders/ Worldfoto / Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
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Paul Souders/ Worldfoto / Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
Clark's anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia.
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David Fleetham / Clark's anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia.
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean.
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David Fleetham / Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean.
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii.
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David Fleetham / Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii.
Blue-headed Wagtail (Motacilla flava) calling, Poland
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Tomasz Zawadzki/ BIA / Blue-headed Wagtail (Motacilla flava) calling, Poland
Great Reed-Warbler (Acrocephalus arundinaceus) calling, Poland
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Tomasz Zawadzki/ BIA / Great Reed-Warbler (Acrocephalus arundinaceus) calling, Poland
Chestnut-bellied Cuckoo (Hyetornis pluvialis) calling, Jamaica
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Glenn Bartley/ BIA / Chestnut-bellied Cuckoo (Hyetornis pluvialis) calling, Jamaica
Patagonian Sierra-Finch (Phrygilus patagonicus) male calling, Andes, Chile
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Glenn Bartley/ BIA / Patagonian Sierra-Finch (Phrygilus patagonicus) male calling, Andes, Chile
Great white shark (Carcharodon carcharias), two below surface. Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias), two below surface. Guadalupe Island, Mexico.
Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico.
Hippopotamus (Hippopotamus amphibius) in territorial display, Mkhuze Game Reserve, KwaZulu-Natal, South Africa
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Winfried Wisniewski / Hippopotamus (Hippopotamus amphibius) in territorial display, Mkhuze Game Reserve, KwaZulu-Natal, South Africa
Hippopotamus (Hippopotamus amphibius) splashing in pool in evening light. Mana Pools National Park, Zimbabwe.
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Tony Heald / Hippopotamus (Hippopotamus amphibius) splashing in pool in evening light. Mana Pools National Park, Zimbabwe.
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May.
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Loic Poidevin / Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May.
Humpback Whale (Megaptera novaeangliae) trio gulp feeding, Inside Passage, southeast Alaska
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Hiroya Minakuchi / Humpback Whale (Megaptera novaeangliae) trio gulp feeding, Inside Passage, southeast Alaska
Hippopotamus (Hippopotamus amphibius) in Chobe River, Botswana.
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Guy Edwardes / Hippopotamus (Hippopotamus amphibius) in Chobe River, Botswana.
Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
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Guy Edwardes / Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
Humpback Whale (Megaptera novaeangliae) pod gulp feeding, Inside Passage, southeast Alaska
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Hiroya Minakuchi / Humpback Whale (Megaptera novaeangliae) pod gulp feeding, Inside Passage, southeast Alaska
Humpback Whale (Megaptera novaeangliae) pod gulp feeding, Inside Passage, southeast Alaska
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Hiroya Minakuchi / Humpback Whale (Megaptera novaeangliae) pod gulp feeding, Inside Passage, southeast Alaska
Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
Black-and-white Warbler (Mniotilta varia) male calling in spring, Maine
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Scott Leslie / Black-and-white Warbler (Mniotilta varia) male calling in spring, Maine
Swallow-tailed Nightjar (Uropsalis segmentata) female in defensive posture on nest, South America
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Murray Cooper / Swallow-tailed Nightjar (Uropsalis segmentata) female in defensive posture on nest, South America
Great Kiskadee (Pitangus sulphuratus) and Green Jay (Cyanocorax yncas) fighting, Texas
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Alan Murphy/ BIA / Great Kiskadee (Pitangus sulphuratus) and Green Jay (Cyanocorax yncas) fighting, Texas
European Bee-eater (Merops apiaster) pair squabbling, Saxony-Anhalt, Germany
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Thomas Hinsche/ BIA / European Bee-eater (Merops apiaster) pair squabbling, Saxony-Anhalt, Germany
Yellow-bellied Marmot (Marmota flaviventris) yawning, central Montana
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Donald M. Jones / Yellow-bellied Marmot (Marmota flaviventris) yawning, central Montana
Elk (Cervus elaphus) bull bugling, western Canada
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Donald M. Jones / Elk (Cervus elaphus) bull bugling, western Canada
Curve-billed Thrasher (Toxostoma curvirostre) feeding on cactus flower nectar, southern Arizona. Sequence 2 of 2
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Donald M. Jones / Curve-billed Thrasher (Toxostoma curvirostre) feeding on cactus flower nectar, southern Arizona. Sequence 2 of 2
Cactus Wren (Campylorhynchus brunneicapillus) calling, southwestern Arizona
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Donald M. Jones / Cactus Wren (Campylorhynchus brunneicapillus) calling, southwestern Arizona
Cactus Wren (Campylorhynchus brunneicapillus) pair with one calling, southwestern Arizona
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Donald M. Jones / Cactus Wren (Campylorhynchus brunneicapillus) pair with one calling, southwestern Arizona
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
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Alex Mustard / Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Saddleback Galapagos Tortoise (Chelonoidis nigra hoodensis) in defensive posture, Espanola Island, Galapagos Islands, Ecuador
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Tui De Roy / Saddleback Galapagos Tortoise (Chelonoidis nigra hoodensis) in defensive posture, Espanola Island, Galapagos Islands, Ecuador
Eastern Meadowlark (Sturnella magna) male calling, Texas
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Alan Murphy/ BIA / Eastern Meadowlark (Sturnella magna) male calling, Texas
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
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Mark Carwardine / Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Grasshopper Warbler (Locustella naevia) calling, Asturias, Spain
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Mario Suarez Porras/ BIA / Grasshopper Warbler (Locustella naevia) calling, Asturias, Spain
Magellanic Penguin (Spheniscus magellanicus) pair calling near chicks, Saunders Island, Falkland Islands
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Tui De Roy / Magellanic Penguin (Spheniscus magellanicus) pair calling near chicks, Saunders Island, Falkland Islands
Southern Elephant Seal (Mirounga leonina) male calling and bleeding after fight, Puerto Madryn, Argentina
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Agustin Esmoris / Southern Elephant Seal (Mirounga leonina) male calling and bleeding after fight, Puerto Madryn, Argentina
Velvet-purple Coronet (Boissonneaua jardini) hummingbird in defensive posture, Las Tangaras Bird Reserve, Colombia
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Thomas Marent / Velvet-purple Coronet (Boissonneaua jardini) hummingbird in defensive posture, Las Tangaras Bird Reserve, Colombia
Black-bellied Whistling Duck (Dendrocygna autumnalis) flying, Florida
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Ron Bielefeld/ BIA / Black-bellied Whistling Duck (Dendrocygna autumnalis) flying, Florida
Black-winged Lapwing (Vanellus melanopterus) calling during defensive display, KwaZulu-Natal, South Africa
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Eric Sohn Joo Tan/ BIA / Black-winged Lapwing (Vanellus melanopterus) calling during defensive display, KwaZulu-Natal, South Africa
Eurasian Wryneck (Jynx torquilla) calling, Aosta Valley, Italy
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Alain Ghignone/ BIA / Eurasian Wryneck (Jynx torquilla) calling, Aosta Valley, Italy
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species.
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Franco Banfi / Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species.
American crocodile (Crocodylus acutus) extreme close up with jaws open, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
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Claudio Contreras / American crocodile (Crocodylus acutus) extreme close up with jaws open, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
RF- Lesser black-backed gulls (Larus fuscus) calling from shoreline, beaks open, England, UK, May. (This image may be licensed either as rights managed or royalty free.)
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Ernie Janes / RF- Lesser black-backed gulls (Larus fuscus) calling from shoreline, beaks open, England, UK, May. (This image may be licensed either as rights managed or royalty free.)
American Alligator (Alligator mississippiensis) split level, Everglades, USA, January.
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Ingo Arndt / American Alligator (Alligator mississippiensis) split level, Everglades, USA, January.
Marsh Wren (Cistothorus palustris) calling, British Columbia, Canada
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Alan Murphy/ BIA / Marsh Wren (Cistothorus palustris) calling, British Columbia, Canada
Hepatic Tanager (Piranga flava) male calling, Arizona
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Alan Murphy/ BIA / Hepatic Tanager (Piranga flava) male calling, Arizona
Golden-fronted Woodpecker (Melanerpes aurifrons) male calling, Texas
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Alan Murphy/ BIA / Golden-fronted Woodpecker (Melanerpes aurifrons) male calling, Texas
Grasshopper Sparrow (Ammodramus savannarum) male calling, Arizona
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Alan Murphy/ BIA / Grasshopper Sparrow (Ammodramus savannarum) male calling, Arizona
Vervet monkey (Chlorocebus pygerythrus) yawning while sitting on a fallen tree in forests, Limpopo River, Kruger National Park, South Africa.
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Neil Aldridge / Vervet monkey (Chlorocebus pygerythrus) yawning while sitting on a fallen tree in forests, Limpopo River, Kruger National Park, South Africa.
Lioness (Panthera leo) yawning in early morning light on Kariega Game Reserve, South Africa.
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Neil Aldridge / Lioness (Panthera leo) yawning in early morning light on Kariega Game Reserve, South Africa.
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean.
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Alex Mustard / Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean.
King Penguin (Aptenodytes patagonicus) chick begging for food, Saunders Island, Falkland Islands
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Heike Odermatt / King Penguin (Aptenodytes patagonicus) chick begging for food, Saunders Island, Falkland Islands
Whale Shark (Rhincodon typus) feeding at Bagan (floating fishing platform),  Cenderawasih Bay, West Papua, Indonesia. Winner of the Man and Nature Portfolio Award in the Terre Sauvage Nature Images Aw...
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Pete Oxford / Whale Shark (Rhincodon typus) feeding at Bagan (floating fishing platform), Cenderawasih Bay, West Papua, Indonesia. Winner of the Man and Nature Portfolio Award in the Terre Sauvage Nature Images Aw...
Jacare / Yacare caiman (Caiman crocodilus yacare) two caymans, one with mouth open, Pantanal, Brazil.
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Bence Mate / Jacare / Yacare caiman (Caiman crocodilus yacare) two caymans, one with mouth open, Pantanal, Brazil.
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.
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Bence Mate / Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.
Cormorants (Phalacrocorax carbo) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
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Bence Mate / Cormorants (Phalacrocorax carbo) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
Grey heron (Ardea cinerea) close up with large fish in beak, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February.
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Bence Mate / Grey heron (Ardea cinerea) close up with large fish in beak, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February.
Collared aracari (Pteroglossus torquatus) two perched, possibly courtship feeding, Costa Rica.
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Bence Mate / Collared aracari (Pteroglossus torquatus) two perched, possibly courtship feeding, Costa Rica.
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.
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Bence Mate / Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.
Great egret (Ardea alba) with green colouring near beak during breeding season, Lake Csaj, Pusztaszer, Hungary, April.
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Bence Mate / Great egret (Ardea alba) with green colouring near beak during breeding season, Lake Csaj, Pusztaszer, Hungary, April.
Grey herons (Ardea cinerea) fighting in water, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.
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Bence Mate / Grey herons (Ardea cinerea) fighting in water, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.
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Bence Mate / Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills whilst feeding, Lake Kerkini, Greece, February.