Anup Shah / African Lion (Panthera leo) juvenile reaching out with curiosity, Masai Mara National Reserve, Kenya. March

Dietmar Nill / Tengmalm's Owl (Aegolius funerus) pouncing on mouse, Germany

Pal Hermansen / Close up of Goshawk (Accipiter gentilis) feathers, with one foot held up, Trondelag, Norway. Winner of the Attention to Detail Category of the Bird Photographer Competition 2019.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) egg hatching on father's feet. Atka Bay, Antarctica. August. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

John Chan / Killdeer (Charadrius vociferus) stamping its feet to create vibrations in the sand as it forages for insects, Southern California, USA, August.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.

Sven Zacek / Eurasian beaver (Castor fiber) on ice besides reeds in Emajogi River, Tartumaa, Southern Estonia. January.

Karine Aigner / Bengal tiger (Panthera tigris tigris) licking paw. Ranthambore National Park, Rajasthan, India.

Tui De Roy / Blue-footed booby (Sula nebouxii) landing. Punta Vicente Roca, Isabela Island, Galapagos.

Tui De Roy / Blue-footed booby (Sula nebouxii) coming in to land. Punta Vicente Roca, Isabela Island, Galapagos.

Phil Savoie / Point-to-Point horse racing, low angle view of racehorse jumping fence, Monmouthshire, Wales, UK. March 2014.

John Waters / Close-up of a Coot (Fulica atra) walking on ice, Greylake RSPB Reserve, Somerset Levels, England, UK, February.