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Kaleidoscope pattern formed from picture of Greenbottle blue tarantula (Chromatopelma cyaneopubescens) - see original image number 01482830 EMBARGOED FOR NAT GEO UNTIL the end of 2015
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Michael D. Kern / Kaleidoscope pattern formed from picture of Greenbottle blue tarantula (Chromatopelma cyaneopubescens) - see original image number 01482830 EMBARGOED FOR NAT GEO UNTIL the end of 2015
Kaleidoscope pattern formed from picture of Johnstons Chameleon (Chamaeleo johnstoni) face and legs. Restricted for Editorial use until December 2015
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Michael D. Kern / Kaleidoscope pattern formed from picture of Johnstons Chameleon (Chamaeleo johnstoni) face and legs. Restricted for Editorial use until December 2015
Close up of Goshawk (Accipiter gentilis) feathers, with one foot held up,  Trondelag, Norway. Winner of the Attention to Detail Category of the Bird Photographer Competition 2019.
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Pal Hermansen / Close up of Goshawk (Accipiter gentilis) feathers, with one foot held up, Trondelag, Norway. Winner of the Attention to Detail Category of the Bird Photographer Competition 2019.
Sea Anemone (Corynactis australis) group, Bicheno, Tasmania, Australia
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Gary Bell/ Oceanwide / Sea Anemone (Corynactis australis) group, Bicheno, Tasmania, Australia
Aerial of patterns of receding waves, Playa del Silencio, Asturias, Spain. April 2018
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Theo Bosboom / Aerial of patterns of receding waves, Playa del Silencio, Asturias, Spain. April 2018
Common limpets (Patella vulgata) hit by a big wave from the Atlantic ocean at the coast of Crohy head, Donegal, Ireland. November 2017
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Theo Bosboom / Common limpets (Patella vulgata) hit by a big wave from the Atlantic ocean at the coast of Crohy head, Donegal, Ireland. November 2017
Winter landscape at Luskentyre beach, Isle of Lewis, Scotland, North Harris in the background. February 2018.
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Theo Bosboom / Winter landscape at Luskentyre beach, Isle of Lewis, Scotland, North Harris in the background. February 2018.
Powerful waves rolling in at the Atlantic coast of Iceland near Dyrholey during a powerful winter storm. February 2016
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Theo Bosboom / Powerful waves rolling in at the Atlantic coast of Iceland near Dyrholey during a powerful winter storm. February 2016
Sand patterns at sunrise, Praia da Adraga, Sintra, Portugal. December 2016
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Theo Bosboom / Sand patterns at sunrise, Praia da Adraga, Sintra, Portugal. December 2016
Patterns on Carrickfinn beach with sand and water mixed with tannin dyed water from peat stream. Donegal, Ireland, November.
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Theo Bosboom / Patterns on Carrickfinn beach with sand and water mixed with tannin dyed water from peat stream. Donegal, Ireland, November.
Patterns of algae on a limestone rock with sand, White sands beach, Causeway coast, Northern Ireland. April 2016.
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Theo Bosboom / Patterns of algae on a limestone rock with sand, White sands beach, Causeway coast, Northern Ireland. April 2016.
Star shaped sand pattern as photographed at low tide at Luskentyre beach, Lewis and Harris, Scotland, UK, October.
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Theo Bosboom / Star shaped sand pattern as photographed at low tide at Luskentyre beach, Lewis and Harris, Scotland, UK, October.
Sand patterns created by receding waves, Quiberon, Brittany, France. August 2017
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Theo Bosboom / Sand patterns created by receding waves, Quiberon, Brittany, France. August 2017
Termite (Macrotermes mossambicus) winged individual on pile of wings shed by other termites, with winged male Carpenter ants (Camponotus). Gorongosa National Park, Mozambique. Highly commended in the...
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Piotr Naskrecki / Termite (Macrotermes mossambicus) winged individual on pile of wings shed by other termites, with winged male Carpenter ants (Camponotus). Gorongosa National Park, Mozambique. Highly commended in the...
Rough-scaled death adder (Acanthophis rugosus) waving tip of tail as a lure to entice prey within striking range. Mount Isa, Queensland Australia.
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Robert Valentic / Rough-scaled death adder (Acanthophis rugosus) waving tip of tail as a lure to entice prey within striking range. Mount Isa, Queensland Australia.
RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image...
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Alex Mustard / RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image...
Amazonian millipede with parasites. Sumaco National Park, Napo, Ecuador
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Lucas Bustamante / Amazonian millipede with parasites. Sumaco National Park, Napo, Ecuador
Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern.
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Ernie Janes / Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern.
Leopard (Panthera pardus) fur, Jao Reserve, Botswana
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Suzi Eszterhas / Leopard (Panthera pardus) fur, Jao Reserve, Botswana
Sword-grass moth (Xylena exsoleta) caterpillar on Field gladiolus (Gladiolus italicus). Cyprus. April.
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Edwin Giesbers / Sword-grass moth (Xylena exsoleta) caterpillar on Field gladiolus (Gladiolus italicus). Cyprus. April.
Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
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Steven David Miller / Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July.
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Marie Read / Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July.
Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
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Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Snowy Owl (Nyctea scandiaca) female perched, Amherst Island, Ontario, Canada, January.
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Marie Read / Snowy Owl (Nyctea scandiaca) female perched, Amherst Island, Ontario, Canada, January.
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions.
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Marie Read / Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions.
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July.
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Marie Read / Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July.
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico.
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Franco Banfi / Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico.
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.
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Franco Banfi / Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.
Mixed coniferous and deciduous forest after snowfall in autumn, Wells Gray Provincial Park, British Columbia, Canada
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Jeff Foott / Mixed coniferous and deciduous forest after snowfall in autumn, Wells Gray Provincial Park, British Columbia, Canada
Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
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Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
Ural owl (Strix uralensis) female stretching wings whilst perched in Spruce (Picea sp) tree. Tartumaa, Southern Estonia. May.
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Sven Zacek / Ural owl (Strix uralensis) female stretching wings whilst perched in Spruce (Picea sp) tree. Tartumaa, Southern Estonia. May.
Beech (Fagus sylvatica) woodland canopy in winter, Cranborne Chase, Dorset, England, UK.
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Guy Edwardes / Beech (Fagus sylvatica) woodland canopy in winter, Cranborne Chase, Dorset, England, UK.
Beech (Fagus sylvatica) woodland canopy in winter, Cranborne Chase, Dorset, England, UK.
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Guy Edwardes / Beech (Fagus sylvatica) woodland canopy in winter, Cranborne Chase, Dorset, England, UK.
Wave in sea off Isle of Harris, Outer Hebrides, Scotland, UK. Long time exposure
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Guy Edwardes / Wave in sea off Isle of Harris, Outer Hebrides, Scotland, UK. Long time exposure
Natural Rock Pattern, Northumberland Coast, England
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Guy Edwardes / Natural Rock Pattern, Northumberland Coast, England
Water flowing over weathered rock with natural patterns, Northumberland Coast, England, UK.
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Guy Edwardes / Water flowing over weathered rock with natural patterns, Northumberland Coast, England, UK.
Close up of a sunflower (Helianthus annuus) head, Provence, France.
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Guy Edwardes / Close up of a sunflower (Helianthus annuus) head, Provence, France.
Forested hills in mist, Ngong Hills, Kenya
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Paul Souders/ Worldfoto / Forested hills in mist, Ngong Hills, Kenya
Bengal tiger (Panthera tigris tigris) licking paw. Ranthambore National Park, Rajasthan, India.
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Karine Aigner / Bengal tiger (Panthera tigris tigris) licking paw. Ranthambore National Park, Rajasthan, India.
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos.
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Nick Hawkins / Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos.
Spoonbill (Platalea leucorodia) preening, close up. Hungary. May.
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Paul Hobson / Spoonbill (Platalea leucorodia) preening, close up. Hungary. May.
Cracked frozen lake following the first snow of winter, aerial view. Akershus, Norway. December 2018.
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Pal Hermansen / Cracked frozen lake following the first snow of winter, aerial view. Akershus, Norway. December 2018.
Skating area labyrinth surrounded by snow, aerial view. Akershus, Norway. December 2017.
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Pal Hermansen / Skating area labyrinth surrounded by snow, aerial view. Akershus, Norway. December 2017.
Guanaco (Lama guanicoe) herd, Torres del Paine National Park, Patagonia, Chile
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Ingo Arndt / Guanaco (Lama guanicoe) herd, Torres del Paine National Park, Patagonia, Chile
Aurora Borealis reflected in Polar Lake, conifers silhouetted at edge. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
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Jack Dykinga / Aurora Borealis reflected in Polar Lake, conifers silhouetted at edge. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Aurora Borealis display above conifers and Polar Lake. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
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Jack Dykinga / Aurora Borealis display above conifers and Polar Lake. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Aurora Borealis reflected in Polar Lake, wetland and conifers silhouetted along shore. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
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Jack Dykinga / Aurora Borealis reflected in Polar Lake, wetland and conifers silhouetted along shore. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Marble formations in alpine landscape. Lahko National Park, Fjell, Norway, July.
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Erlend Haarberg / Marble formations in alpine landscape. Lahko National Park, Fjell, Norway, July.
Clark's anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia.
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David Fleetham / Clark's anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia.
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines.
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David Fleetham / Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines.
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean.
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David Fleetham / Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean.
Red-billed Oxpecker (Buphagus erythrorhynchus) on Burchell's Zebra (Equus burchellii), Mkhuze Game Reserve, KwaZulu-Natal, South Africa
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Richard Du Toit / Red-billed Oxpecker (Buphagus erythrorhynchus) on Burchell's Zebra (Equus burchellii), Mkhuze Game Reserve, KwaZulu-Natal, South Africa
Day / Big blue octopus (Octopus cyanea) with diver in background. Hawaii. September 2012.
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David Fleetham / Day / Big blue octopus (Octopus cyanea) with diver in background. Hawaii. September 2012.
Whale shark (Rhincodon typus), a commercial encounter for tourists with feeder above on canoe. Oslob, Cebu, Philippines. April 2017.
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David Fleetham / Whale shark (Rhincodon typus), a commercial encounter for tourists with feeder above on canoe. Oslob, Cebu, Philippines. April 2017.
Sumatran tiger (Panthera tigris sumatrae) close up portrait, captive.
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Paul Williams / Sumatran tiger (Panthera tigris sumatrae) close up portrait, captive.
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017.
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Nick Garbutt / Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017.
Seaweed in tide pool, Mangurstadh Beach, Isle of Lewis, Outer Hebrides, October. Highly commended in the Creative Visions category of Wildlife photographer of the year Awards (WPOY) Competition 2018.
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Theo Bosboom / Seaweed in tide pool, Mangurstadh Beach, Isle of Lewis, Outer Hebrides, October. Highly commended in the Creative Visions category of Wildlife photographer of the year Awards (WPOY) Competition 2018.
Landscape in Fjallabak Nature Reserve, with circular patterns on the ground, Iceland, September 2017. Highly commended in the Landscape category of the GDT European Wildlife Photographer of the Year A...
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Theo Bosboom / Landscape in Fjallabak Nature Reserve, with circular patterns on the ground, Iceland, September 2017. Highly commended in the Landscape category of the GDT European Wildlife Photographer of the Year A...
Large flock of Red knot (Calidris canutus) moving at high tide roost, in winter plumage, Snettisham RSPB Reserve, Norfolk, England, UK, September.
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Chris Gomersall / Large flock of Red knot (Calidris canutus) moving at high tide roost, in winter plumage, Snettisham RSPB Reserve, Norfolk, England, UK, September.
Saddle-billed stork (Ephippiorhynchus senegalensis), Moremi Game Reserve, Botswana.
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Guy Edwardes / Saddle-billed stork (Ephippiorhynchus senegalensis), Moremi Game Reserve, Botswana.
Trees with new foliage, view from Eggardon Hill, Bridport, Dorset, England, UK. May 2010.
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Guy Edwardes / Trees with new foliage, view from Eggardon Hill, Bridport, Dorset, England, UK. May 2010.
Rock formations at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
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Guy Edwardes / Rock formations at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
Rocky shore at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
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Guy Edwardes / Rocky shore at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
Rocky shore at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
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Guy Edwardes / Rocky shore at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
Clavell Tower, Kimmeridge Bay, Jurrasic Coast, Isle of Purbeck, Dorset, England, UK. November2011.
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Guy Edwardes / Clavell Tower, Kimmeridge Bay, Jurrasic Coast, Isle of Purbeck, Dorset, England, UK. November2011.
Warbeth Beach, Stromness, Orkney with view to Hoy. Orkney Islands, Scotland, UK. August 2014.
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Guy Edwardes / Warbeth Beach, Stromness, Orkney with view to Hoy. Orkney Islands, Scotland, UK. August 2014.
Rock formations at Kimmeridge Bay, Isle of Purbeck, Dorset, England, UK. October 2011.
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Guy Edwardes / Rock formations at Kimmeridge Bay, Isle of Purbeck, Dorset, England, UK. October 2011.
Saltmarsh on the Isle of Harris, Outer Hebrides, Scotland, UK. November 2016.
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Guy Edwardes / Saltmarsh on the Isle of Harris, Outer Hebrides, Scotland, UK. November 2016.
Saltmarsh. Isle of Harris, Scotland, UK. March, 2015.
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Guy Edwardes / Saltmarsh. Isle of Harris, Scotland, UK. March, 2015.
Oldworld Swallowtail (Papilio machaon) wing, Baden-Wurttemberg, Germany
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Ralph Martin/ BIA / Oldworld Swallowtail (Papilio machaon) wing, Baden-Wurttemberg, Germany
Frozen water patterns in Klein Schietveld, Brasschaat, Belgium
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Bernard Castelein / Frozen water patterns in Klein Schietveld, Brasschaat, Belgium
Close up of the pattern on the Hawaiian spotted flatworm (Pseudobiceros)  endemic to Hawaii.
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David Fleetham / Close up of the pattern on the Hawaiian spotted flatworm (Pseudobiceros) endemic to Hawaii.
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean.
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Alex Mustard / Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean.
Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean.
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Alex Mustard / Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean.
RF -  Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean. (This image may be licensed either as rights managed or royalty free.)
Mountains along lake with choco rainforest, Utria National Park, Colombia
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Chien Lee / Mountains along lake with choco rainforest, Utria National Park, Colombia
Glacier, Breidamerkurjokull, Jokalsarlon Lagoon, Iceland
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Yva Momatiuk & John Eastcott / Glacier, Breidamerkurjokull, Jokalsarlon Lagoon, Iceland
American crocodiles (Crocodylus acutus) in the Rio Tarcoles, Costa Rica.
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Jen Guyton / American crocodiles (Crocodylus acutus) in the Rio Tarcoles, Costa Rica.
School of Longfin spadefish (Platax teira), Kimbe Bay, West New Britain, Papua New Guinea
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Bert Willaert / School of Longfin spadefish (Platax teira), Kimbe Bay, West New Britain, Papua New Guinea
Limestone pinnacles, Tsingy de Bemaraha National Park, Madagascar
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Chien Lee / Limestone pinnacles, Tsingy de Bemaraha National Park, Madagascar
Rainforest at sunrise, Sani Lodge, Ecuador
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Paul Bertner / Rainforest at sunrise, Sani Lodge, Ecuador
Forest-covered hills and river, Taironaka Lodge, Colombia
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Paul Bertner / Forest-covered hills and river, Taironaka Lodge, Colombia
Maple (Acer sp) leaves and needles in autumn, Acadia National Park, Maine
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Jeff Foott / Maple (Acer sp) leaves and needles in autumn, Acadia National Park, Maine
Deciduous forest and pond in autumn, Jordan Pond, Mount Desert Island, Acadia National Park, Maine
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Jeff Foott / Deciduous forest and pond in autumn, Jordan Pond, Mount Desert Island, Acadia National Park, Maine
Deciduous forest in autumn, Acadia National Park, Maine
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Jeff Foott / Deciduous forest in autumn, Acadia National Park, Maine
Deciduous forest in autumn, Acadia National Park, Maine
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Jeff Foott / Deciduous forest in autumn, Acadia National Park, Maine
Aerial shot of patterns in a frozen lake, Lake Langen, Akershus, Norway, December 2017.
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Pal Hermansen / Aerial shot of patterns in a frozen lake, Lake Langen, Akershus, Norway, December 2017.
Aerial shot of field with tractor tracks, Ski, Akershus, Norway, November 2017.
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Pal Hermansen / Aerial shot of field with tractor tracks, Ski, Akershus, Norway, November 2017.
Aerial shot of an ice skating arena, Lake Langen, Akershus, Norway, December 2017.
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Pal Hermansen / Aerial shot of an ice skating arena, Lake Langen, Akershus, Norway, December 2017.
Aerial shot of lake covered by the first snow, Bindingsvann, Akershus, November Norway, November 2017.
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Pal Hermansen / Aerial shot of lake covered by the first snow, Bindingsvann, Akershus, November Norway, November 2017.
Garden tiger moth (Arctia caja) Killard Point NNR, Ballyhornan, County Down, Northern Ireland.July.
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Robert Thompson / Garden tiger moth (Arctia caja) Killard Point NNR, Ballyhornan, County Down, Northern Ireland.July.
Floating sweet-grass (Glyceria fluitans) in lake, Bjornlandet National Park, North Sweden. Finalist in the Plants and Fungi category of the Wildlife Photographer of the Year Awards (WPOY) Competition...
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Theo Bosboom / Floating sweet-grass (Glyceria fluitans) in lake, Bjornlandet National Park, North Sweden. Finalist in the Plants and Fungi category of the Wildlife Photographer of the Year Awards (WPOY) Competition...
Crustose lichens (Lecanorales) on rocks in Swedish Padjelanta National Park, Laponia World Heritage Site, Sweden. August.  Highly commended in the GDT  European Wildlife Photographer of the Year  com...
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Erlend Haarberg / Crustose lichens (Lecanorales) on rocks in Swedish Padjelanta National Park, Laponia World Heritage Site, Sweden. August. Highly commended in the GDT European Wildlife Photographer of the Year com...
Two small pearl-bordered fritillary butterflies (Boloria selene) resting on bluebell, Marsland mouth, North Devon, UK. May 2017.
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Ross Hoddinott / Two small pearl-bordered fritillary butterflies (Boloria selene) resting on bluebell, Marsland mouth, North Devon, UK. May 2017.
Agave (Agave utahensis) close up of spines with Pinyon pine cone (Pinus edulis) wedged amongst them. Grand Canyon-Parashant National Monument, Arizona, USA
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Jack Dykinga / Agave (Agave utahensis) close up of spines with Pinyon pine cone (Pinus edulis) wedged amongst them. Grand Canyon-Parashant National Monument, Arizona, USA
Blue-legged Chameleon (Calumma crypticum) curled tail, Ranomafana National Park, Madagascar
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Paul Bertner / Blue-legged Chameleon (Calumma crypticum) curled tail, Ranomafana National Park, Madagascar