Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Andy Rouse / Little Bittern (Ixobrychus minutus) hunting in reeds, Bulgaria. Horizontal version of 01609394. Highly honoured in the Bird Category of the Nature's Best Photography Competition 2019.

Andy Rouse / African lion (Panthera leo) female carrying young cub, Masai Mara, Kenya, Africa. Highly commended in the African Wildlife Category of the Nature's Best Photography Competition 2019.

Diane McAllister / Polar bears (Ursus maritimus) fighting in water. Beaufort Sea , Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Nature's Best Photography Competition 2019.

Sven Zacek / Aerial view of a small forest lake in Karula National Park, Valgamaa county, Southern Estonia. Highly commended in the Wildlife Photographer of the Year Awards 2019.

Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) two adults brooding chicks age 5 weeks, Antarctica. Highly commended in the Polar Passion Category of the Nature's Best Photography Awards 2019.

Claudio Contreras / Californian sea lion (Zalophus californianus) swimming in Southern sea palm (Eisenia arborea) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphe...

Kevin Schafer / Bumblebee (Bombus sp) on flower, Juneau, Alaska

Chase Dekker / Humpback Whale (Megaptera novaeangliae) pod gulp feeding on Northern Anchovy (Engraulis mordax), Monterey Bay, California

Chase Dekker / Humpback Whale (Megaptera novaeangliae) calf breaching, Monterey Bay, California

Chase Dekker / Humpback Whale (Megaptera novaeangliae) calf breaching, Monterey Bay, California

Chase Dekker / Humpback Whale (Megaptera novaeangliae) calf breaching, Monterey Bay, California

Chase Dekker / Humpback Whale (Megaptera novaeangliae) breaching at sunset, Monterey Bay, California

Chase Dekker / African Lion (Panthera leo) male, Masai Mara, Kenya

Chase Dekker / African Lion (Panthera leo) cub, Masai Mara, Kenya

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) wading, Amboseli National Park, Kenya

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya

Chase Dekker / Humpback Whale (Megaptera novaeangliae) breaching, Monterey Bay, California

Gerry Ellis / Slash-and-burn deforestation for farming, near Mefou Primate Sanctuary, Cameroon

Gerry Ellis / Western Lowland Gorilla (Gorilla gorilla gorilla) silverback, Limbe Wildlife Centre, Cameroon

Gerry Ellis / Chimpanzee (Pan troglodytes) orphan Larry playing in tree, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Gerry Ellis / Chimpanzee (Pan troglodytes) orphans, Larry investigating Daphne's tongue, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Gerry Ellis / Chimpanzee (Pan troglodytes) orphan Larry, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Gerry Ellis / Chimpanzee (Pan troglodytes) orphan Larry playing in forest nursery, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Gerry Ellis / Chimpanzee (Pan troglodytes) orphan Larry scratching, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Winfried Wisniewski / White Stork (Ciconia ciconia) on nest, Extremadura, Spain

Jami Tarris / Polar Bear (Ursus maritimus) cooling off, Spitsbergen, Svalbard, Norway

Jami Tarris / Walrus (Odobenus rosmarus) bull, Spitsbergen, Svalbard, Norway

Jami Tarris / Polar Bear (Ursus maritimus) hunting Bearded Seal (Erignathus barbatus) pup, Spitsbergen, Svalbard, Norway

Jami Tarris / Polar Bear (Ursus maritimus) on ice floe, Spitsbergen, Svalbard, Norway

Jami Tarris / Polar Bear (Ursus maritimus) on shore, Spitsbergen, Svalbard, Norway

Jami Tarris / Arctic Fox (Alopex lagopus) running in snow, Spitsbergen, Svalbard, Norway

Jami Tarris / Golden Eagle (Aquila chrysaetos) huntress, Altai Mountains, Bayan Ulgii, Mongolia

Jami Tarris / Caribou (Rangifer tarandus) entering teepee for food, Khovsgol, Mongolia

Juergen & Christine Sohns / African Lion (Panthera leo) six week old cubs, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / African Lion (Panthera leo) six week old cub, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / African Lion (Panthera leo) male drinking at waterhole, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / African Lion (Panthera leo) male, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / African Lion (Panthera leo) juvenile male, Mountain Zebra National Park, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pregnant female and juveniles on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pregnant female and juveniles on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pair on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pair on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pair on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) pair on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) on alert, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Meerkat (Suricata suricatta) scratching itself, Tswalu Game Reserve, South Africa

Juergen & Christine Sohns / Mountain Zebra (Equus zebra), Mountain Zebra National Park, South Africa

Juergen & Christine Sohns / African Wild Dog (Lycaon pictus) group attacking Cape Warthog (Phacochoerus aethiopicus), Tswalu Game Reserve, South Africa

Anup Shah / Olive Baboon (Papio anubis) male with three month old young, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) fourteen year old sub-adult male, named Fundi, making submissive facial gesture, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-three year old female, named Gremlin, scratching face, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty-five year old female, named Nasa, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) four year old infant male, named Gizmo, foraging for insects in log, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty year old male, named Titan, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) seven year old juvenile male, named Siri, feeding on vines, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty-five year old female, named Tanga, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) four year old infant male, named Gizmo, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) eighteen year old male, named Sampson, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) four year old infant male, named Gizmo, Gombe National Park, Tanzania

Richard Du Toit / Greater Double-collared Sunbird (Nectarinia afra) male on Pincushion (Leucospermum sp) flower, Western Cape, South Africa

Heike Odermatt / Walrus (Odobenus rosmarus), Hinlopen Strait, Svalbard, Norway

Heike Odermatt / Walrus (Odobenus rosmarus), Svalbard, Norway

Heike Odermatt / Walrus (Odobenus rosmarus) on ice floe, Hinlopen Strait, Svalbard, Norway

Heike Odermatt / Walrus (Odobenus rosmarus) on ice floe, Hinlopen Strait, Svalbard, Norway

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Heike Odermatt / Walrus (Odobenus rosmarus) on ice floe, Svalbard, Norway

Heike Odermatt / Walrus (Odobenus rosmarus) climbing on ice floe, Svalbard, Norway

Heike Odermatt / Bearded Seal (Erignathus barbatus) on ice floe, Magdalenefjorden, Svalbard, Norway

Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 2 of 3

Martin Willis / Splendid Fairywren (Malurus splendens) female and male, Queensland, Australia

Martin Willis / Red-collared Lorikeet (Trichoglossus rubritorquis) feeding on flower nectar, Kimberley, Western Australia, Australia

Martin Willis / Rainbow Lorikeet (Trichoglossus haematodus) pair preening during courtship, Magnetic Island, Queensland, Australia

Martin Willis / Spinifex Grass (Spinifex sp) in desert, Kata Tjuta, Australia

Martin Willis / Mangroves along coast and boat, Deluge Inlet, Hinchinbrook Island National Park, Queensland, Australia

Martin Willis / Budgerigar (Melopsittacus undulatus) father with begging chick at nest cavity, Queensland, Australia

Suzi Eszterhas / Golden Lion Tamarin (Leontopithecus rosalia), Atlantic Forest, Brazil

Suzi Eszterhas / Golden Lion Tamarin (Leontopithecus rosalia), Atlantic Forest, Brazil

Suzi Eszterhas / Golden Lion Tamarin (Leontopithecus rosalia), Atlantic Forest, Brazil

Ann & Steve Toon / Meerkats (Suricata suricatta) on the move, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January. Second place in the Mammals Category of the GDT European Nature Photographer of the Year...

Gary Bell/ Oceanwide / Australian Pelican (Pelecanus conspicillatus) flock, New South Wales, Australia

Gary Bell/ Oceanwide / Spotted Wobbegong (Orectolobus maculatus), Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Port Jackson Shark (Heterodontus portusjacksoni), Australia

Gary Bell/ Oceanwide / Grey Nurse Shark (Carcharias taurus), Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Tropical island, Keeling Islands, Australia

Gary Bell/ Oceanwide / Tropical island, Keeling Islands, Australia

Gary Bell/ Oceanwide / Tropical island, Keeling Islands, Australia

Gary Bell/ Oceanwide / Tropical island, Keeling Islands, Australia

Gary Bell/ Oceanwide / Surfer, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Lichen (Caloplaca sp) covered coastal rock, Flinders Island, Tasmania, Australia

Gary Bell/ Oceanwide / Coastal rock arch at sunset, Bermagui, New South Wales, Australia

Gary Bell/ Oceanwide / Cave along coast, Loch Ard Gorge, Port Campbell National Park, Victoria, Australia

Gary Bell/ Oceanwide / Granite rocks with lichen and Milky Way, Bicheno, Tasmania, Australia