Sandesh Kadur / Female Dussumier's malabar langur (Semnopithecus dussumieri) feeding with baby, picks it up and walks away, Ranthambore National Park, Rajasthan, India. 2016.

Bence Mate / Great egret (Ardea alba) hunting fish, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.

Tony Heald / Asian elephant (Elephas maximus) domesticated elephant working, picking reeds, Kaziranga National Park, Assam, India, March 2009.

Cyril Ruoso / African elephant (Loxodonta africana) hanging upside down, during translocation by Kenya Wildlife Service, due to over population, Mwaluganje Reserve, Kenya. October 2006.

Espen Bergersen / Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March

Chris & Monique Fallows / Orca male (Orcinus orca) raising pectoral flipper out of the water, Prince William Sound, Alaska, USA.

Chris Mattison,Chris Mattison / Rubber Boa (Charina bottae) in defensive posture, raising tail to mimic head. California, USA, November.

Anup Shah / Spotted hyena (Crocuta crocuta) preparing to carry a pup aged 2-3 months. Masai Mara National Reserve, Kenya, July

Tony Heald / Man of the Toda tribe lifting a heavy stone to show he is ready for marriage. Nilgiri Mountains, Tamil Nadu, India. 2014.

Heather Angel / Bee beetle (Trichius fasciatus), a bee mimic, nectaring on Umbel-flowered catchfly (Silene compacta). Picks up pollen on hairy body. Russian Caucasus. June.

SCOTLAND: The Big Picture / Volunteers working in Trees for Life's native tree nursery, Dundreggan Conservation Estate, Dumfries and Galloway, Scotland, UK. .

Dave Bevan / Pair of Slow worms (Anguis fragilis) exposed when roofing felt is lifted, Carmarthenshire, Wales, UK, May.