Andy Rouse / African elephant (Loxodonta africana) on plains under stormy sky, Marakele National Park, Waterberg Biosphere, South Africa

John Shaw / Shoreline of Lake Powell, seen from Alstrom Point on Romana Mesa. Glen Canyon National Recreation Area, on the Arizona/Utah border, USA, January.

Jack Dykinga / Morning light with autumn coloured Cottonwoods (Populus sp) trees lining the river. Valley of the Gods, Colorado Plateau, of the Great Basin Desert, Utah, USA, November 2018.

Anup Shah / Mixed herd of Common or Plains zebra (Equus quagga burchellii) and Eastern white-bearded wildebeest (Connochaetes taurinus) - remote camera. Masai Mara National Reserve, Kenya.

Richard Du Toit / Red-billed Oxpecker (Buphagus erythrorhynchus) on Burchell's Zebra (Equus burchellii), Mkhuze Game Reserve, KwaZulu-Natal, South Africa

Fiona Rogers / Hanuman Langur (Semnopithecus entellus) group on temple, Virupaksha Temple, Hampi, Karnataka, India

Richard Du Toit / Burchell's Zebra (Equus burchellii) trio nuzzling, Itala Game Reserve, South Africa

Bence Mate / Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / African elephant (Loxodonta africana) and Zebra (Equus quagga) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Christophe Courteau / Plains zebra (Equus burchelli) drinking in a dust storm, Boteti River, Kalahari Desert, Botswana.

Sandesh Kadur / Female Dussumier's malabar langur (Semnopithecus dussumieri) feeding with baby, picks it up and walks away, Ranthambore National Park, Rajasthan, India. 2016.

Frederique Olivier / Young Burchell's zebra (Equus burchellii) playing, biting and kicking its mother, Serengeti NP, Tanzania.