Alex Mustard / Portrait of a large female Reef manta ray (Mobula alfredi) swimming over a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Portrait of a Starry grouper (Epinephelus labriformis) over coral. La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.

Alex Mustard / Long exposure portrait of a Tawny nurse shark (Nebris ferrungineus), with Pink whipray (Himantura fai) and Bluefin trevally (Caranx melampygus) at dusk. Vavuu Atoll, Maldives. Indian Ocean

Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the packground sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...

Alex Mustard / Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a colourful nudibranch (Chromodoris annae). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Staffan Widstrand / Painted stork (Mycteria leucocephala) head and beak portrait, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian / Blue peafowl (Pavo cristatus) head portrait, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Gaur or Indian bison (Bos gaurus / Bos frontalis) Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Young Black or Cinereous vulture (Aegypius monachus) Gredos Mountains, Castilla La Mancha, Spain

Staffan Widstrand / White rhinoceros (Ceratothorium simum) Marataba Private Reserve, Marakele National Park, Limpopo, South Africa

Staffan Widstrand / Hawaiian Goose, or Nene (Branta sandvicensis) Maui, Hawaii. Endemic.

Staffan Widstrand / Hawaiian Goose, or Nene (Branta sandvicensis) Maui, Hawaii. Endemic

Robert Valentic / Heath or Littlejohn's tree frog (Litoria littlejohni) from riparian heath in the Parma Creek Nature Reserve west of Jervis Bay, New South Wales, Australia.

Robert Valentic / Eastern Spiny-tailed Gecko (Strophurus williamsi) male, near St George, Queensland, Australia.

Robert Valentic / Cascade Tree Frog (Litoria pearsoniana) in rainforest near Peachester, Sunshine Coast hinterland, Queensland, Australia.

Robert Valentic / Spotted tree frog (Litoria spenceri) from Still Creek in north-eastern Victoria, Australia.

Robert Valentic / Southern knob-tailed gecko (Nephrurus w. wheeleri) male, Cue, southern central Western Australia. January.

Robert Valentic / Midline knob-tailed gecko (Nephrurus vertebralis) from Cue in the central Western Australia, January.

Robert Valentic / Bana bubble-nest frog (Kurixalus banaensis) from high elevation rainforest along the summit trail in Bach Ma National Park, central Vietnam. Dry season.

Robert Valentic / Rough-scaled death adder (Acanthophis rugosus) waving tip of tail as a lure to entice prey within striking range. Mount Isa, Queensland Australia.

Robert Valentic / Vogel's pit viper (Trimeresurus vogeli) male in elevated rainforest habitat within the Bach Ma National Park in central Vietnam. Dry season.

Robert Valentic / Narrow-mouthed frog (Microhyla marmorata) male, from montane rainforest at Phong Lan, Bach Ma National Park, Vietnam. Dry season.

Robert Valentic / Emma gray's forest dragon (Calotes emma emma) female from the botanical garden trail in karst rainforest habitat within the Phong Nha Ke Bang National Park in central Vietnam. Dry season.

Robert Valentic / Brown pricklenape (Acanthosaura lepidogaster) female, Phong Nha-Ke Bang National Park in central Vietnam. Dry season.

Tony Wu / Fringed blenny (Chirolophis japonicus) guarding its eggs. Miyagi Prefecture, Japan.

Alex Mustard / RF - Portrait of Coral hermit crab (Paguritta harmsi) living in a hard coral colony. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)

Alex Mustard / RF - Portrait of a large female Reef manta ray (Mobula alfredi) swimming over a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image may be licens...

Alex Mustard / Portrait of a Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a Blue ringed octopus (Hapalochlaena lunulata) on an algae covered rock. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a Giant frogfish (Antennarius commersoni) on a large Yellow elephant ear sponge (Ianthella basta). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a hidden Corallimorph shrimp (Pliopontonia furtiva) on a coral reef. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Plain spot octopus (Octopus exannulatus) on the seabed at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.

Alex Mustard / Portrait of a Whip coral shrimp (Pontonides ankeri). This species, although widespread and reasonably common, was formally described only in 2007, by Russian zoologist Ivan Marin. Seraya, Bali, Indone...

Alex Mustard / Portrait of a male short snouted seahorse (Hippocampus hippocampus) in sea oak seaweed (Halidrys siliquosa). Devon, England, United Kingdom. British Isles. English Channel. North East Atlantic Ocean.

Lucas Bustamante / Gold-nugget treefrog (Boana picturata) Mashpi, Pichincha, Ecuador

Lucas Bustamante / Black Caiman (Melanosuchus niger) Yasuni National Park, Orellana, Ecuador September 2018

Robin Chittenden / Whooper swan (Cygnus cygnus) Norfolk, England, UK January.

Robin Chittenden / Snow bunting (Plectrophenax nivalis) Norfolk, England, UK, January.

Robin Chittenden / Whooper swan (Cygnus cygnus) Norfolk, England, UK, January.

Lucas Bustamante / Tropical shield mantis (Choeradodis sp.) Sumaco National Park, Napo, Ecuador

Lucas Bustamante / Pinzon Giant-Tortoise (Chelonoidis duncanensis) Pinzon or Duncan Island, Galapagos Islands.

Lucas Bustamante / Green-crowned woodnymph (Thalaurania fannyi) Mashpi, Pichincha, Ecuador

Lucas Bustamante / Sierra Negra giant tortoise (Chelonoidis guntheri) Isabela Island, Galapagos Islands.

Lucas Bustamante / Crested hawk-eagle (Nisaetus cirrhatus) Yala National Park, Southern Province, Sri Lanka

Lucas Bustamante / Scarlet macaw (Ara macao) Tambopata National Reserve, Madre de Dios, Peru.

Lucas Bustamante / Rose-ringed parakeet (Psittacula krameri) Yala National Park, Southern Province, Sri Lanka

Fabio Pupin / Tenerife gecko, (Tarentola delalandii), endemic, Tenerife, Canary Islands, Spain, April.

Franco Banfi / Sperm whale, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean.

Ernie Janes / Celebes crested macaque (Macaca nigra) infant, captive.

Ernie Janes / Blue-eyed black lemur (Eulemur flavifrons) male, captive, occurs in Sulawesi. With digitally added leaves.

Ernie Janes / Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern.

Ernie Janes / Black howler (Alouatta caraya) male female pair. Captive, occurs in South America, with digitally added leaf pattern.

Ernie Janes / Amur leopard (Panthera pardus orientalis) captive, occurs in northern China and Russia. Digitally added shadows.

Ernie Janes / RF - Amur leopard (Panthera pardus orientalis) captive, occurs in northern China and Russia, with digitally added leaves. (This image may be licensed either as rights managed or royalty free.)

Ernie Janes / RF - White-throated toucan (Ramphastos tucanus) captive, occurs in South America, with digitally added leaves. (This image may be licensed either as rights managed or royalty free.)

Ernie Janes / Victoria crowned pigeon (Goura victoria) portrait. Captive, occurs in New Guinea.

Ernie Janes / Bald eagle (Haliaeetus leucocephalus) portrait, captive, occurs in North America.

Ernie Janes / RF - Bald eagle (Haliaeetus leucocephalus) portrait, captive, occurs in North America. (This image may be licensed either as rights managed or royalty free.)

Ernie Janes / Cougar (Puma concolor) portrait, captive, occurs in the Americas. With digitally added leaves.

Ernie Janes / Coppery titi monkey (Callicebus cupreus) portrait, captive, occurs in South America. with digitally added leaves.

Ernie Janes / Common marmoset (Callithrix jacchus) portrait, captive, endemic to Brazil. With digitally added leaves.

Ernie Janes / RF - Celebes crested macaque (Macaca nigra) captive, with digitally added leaf pattern. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / Photographers Steve and Ann Toon, looking through window in Tankwa Karoo National Park, South Africa

Nigel Cattlin / White huacaya alpaca with head up and teeth bared hair growing back after shearing, Berkshire, May

David Woodfall / Red Deer (Cervus elaphus) portrait of head of stag and antlers. Alladale Estate, Scotland, UK, October.

David Woodfall / Lapwing (Vanellus vanellus) looking for food on machir marsh , threatened wader of machir habitat .

David Woodfall / Hedgehog (Erinaceus europaeus )about to enter hedgehog trap put there by Scottish Natural Heritage scientists to catch and remove introduced mammal from The Uists (where they present a threat to Littl...

David Woodfall / Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October.

David Woodfall / Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October.

David Woodfall / Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October.

John Shaw / Ocelot (Leopardus pardalis) hiding in a tree. Porto Jofre, Pantanal, Brazil. September.

John Shaw / Magellanic snipe (Gallinago magellanica magellanica). Sealion Island, Falklands. November.

John Shaw / Yacare caiman (Caiman yacare) Cuiaba River, Pantanal, Brazil, August.

Shane Gross / Portrait of a Thorny seahorse (Hippocampus histrix) near Malapascua, Philippines.

Shane Gross / Head portrait of a Shaws seasnake (Lapemis curtus) underwater near Malapascua, Philippines.

Shane Gross / Portrait of an Ornate ghost pipefish (Solenostomus paradoxus) Anilao, Philippines.

Shane Gross / Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines.

Andy Rouse / European Hare (Lepus europaeus) in grassland. UK. May.

Andy Rouse / European Hare (Lepus europaeus) portrait. UK. May.

Espen Bergersen / Killer whale / orca (Orcinus orca). Spyhopping very close, showing eye. Kvanangen, Troms, Norway. November

Phil Savoie / Eurasian otter (Lutra lutra) portrait, captive. . May

Georgette Douwma / Whitemouth moray eel (Gymnothorax meleagris). Lembeh Strait, North Sulawesi, Indonesia. Indo-Pacific.

Georgette Douwma / Thornback cowfish (Lactoria fornasini). Lembeh Strait, North Sulawesi, Indonesia.

Edwin Giesbers / Lemon-yellow tree frog (Hyla savignyi) looking at camera, portrait. Cyprus. April.

Andy Rouse / Indri Lemur, (Indri indri), in rainforest, Perinet, Madagascar

Andy Rouse / African Lion, (Panthera leo), female hunting, Masai Mara, Kenya, Africa,

Andy Rouse / Galapagos sea lion, (Zalophus wollebaeki), pup on shore, Galapagos.

Andy Rouse / Galapagos dove , (Zenaida galapagoensis), portrait , Santa Fe Island, Galapagos

Andy Rouse / Red fox, (Vulpes vulpes), in bluebells, UK

Andy Rouse / Sally lightfoot crab, (Grapsus grapsus), at sunrise, Galapagos

Andy Rouse / RF - Marine Iguana, (Amblyrhynchus cristatus), warming at sunrise , Fernandina Island, Galapagos.