Lucas Bustamante / Sri Lankan elephant (Elephas maximus maximus) walking past tourist jeep with tourists, Yala National Park, Southern Province, Sri Lanka

Suzi Eszterhas / Leopard (Panthera pardus) five-week-old cub playing with mother, Jao Reserve, Botswana

David Woodfall / European beaver (Castor fiber) rear view in wetlands, in captive breeding programme .Bevis Trust, Carmarthenshire, Wales, UK, May.

Shane P. White / Coconut Crab (Birgus latro) on palm tree, Espiritu Santo, Vanuatu

Jeff Foott / Hiker near waterfall and rainbow, Nevada Fall, Yosemite National Park, California

Paul Hobson / Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October.

Brian Bevan / Pantaloon bee (Dasypoda hurtles) entering burrow with pollen, Bedfordshire, England, UK, August.

Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) silverback sitting on boundary wall between Volcanoes National Park and a Potato crop, looking into valley. Area to be restored to forest. Rwanda.

Tim Fitzharris / Ruby-throated Hummingbird (Archilochus colubris) female and Bumblebee (Bombus sp) flying, Arkansas

Suzi Eszterhas / African Elephant (Loxodonta africana) in defensive posture towards safari vehicle, Sabi Sands Private Game Reserve, South Africa

Graeme Guy/ BIA / Chestnut-headed Bee-eater (Merops leschenaulti) trio with insect prey, Penang, Malaysia

Piotr Naskrecki / Large Striped Swordtail (Graphium antheus) butterflies, Gorongosa National Park, Mozambique