Chris Mattison / Green wasp (Ropalidia sp), two, one guarding nest. Andasibe-Mantadia National Park, Madagascar.

Chris Mattison / Green wasp (Ropalidia sp), two, one guarding nest. Andasibe-Mantadia National Park, Madagascar.

Paul Bertner / Crab Spider (Thomisidae) with Paper Wasp (Ropalidia sp) prey, Antananarivo, Madagascar

Paul Bertner / Paper Wasp (Ropalidia sp) colony, Ankarafantsika National Park, Madagascar

Paul Bertner / Paper Wasp (Ropalidia sp) queen tending to brood, Antananarivo, Madagascar

Paul Bertner / Paper Wasp (Ropalidia sp) group on nest, Ankarafantsika National Park, Madagascar

Paul Bertner / Paper Wasp (Ropalidia sp) queen at nest, Ranomafana National Park, Madagascar

John Abbott / Polistes sp. (Paper Wasp) in flight, captured using high speed flashes Travis County, Texas, USA. Controlled conditions. March

Nick Garbutt / Social Green paper wasp (Ropalidia sp) on paper nest. Ranomafana National Park, SE Madagascar.

Kim Taylor / Paper wasps' nest {Ropalidia sp}, Northern Australia.

Jiri Lochman / Paper wasp (Ropalidia sp.) worker at the nest feeding larvae, Kununurra, Kimberley Region, Western Australia.

Jiri Lochman / Paper wasp (Ropalidia sp.) workers at nest taking care of the brood, Kununurra, Kimberley Region, Western Australia.

Christian Ziegler / Paper Wasp (Ropalidia sp) at nest, Ranomafana National Park, Madagascar