Tom Vezo / Tufted puffins {Lunda cirrhata} fighting, Alaska. Saint Paul Island

Tom Vezo / Tufted puffin {Lunda cirrhata} pair perched on cliff edge, Alaska. Saint Paul Island

Tom Vezo / Tufted puffins {Lunda cirrhata} billing, Alaska. Saint Paul Island

Tom Vezo / Tufted puffin {Lunda cirrhata} perched on cliff edge, Alaska. Saint Paul Island

Bence Mate / Puffin portrait {Fratercula arctica} soft focus, Runde, Norway

Bence Mate / Puffin portrait {Fratercula arctica} Runde, Norway

Steve Knell / Black Guillemot (Cepphus grylle) on lichen-covered rock with small plaice in its beak, UK

Steve Knell / Black Guillemot (Cepphus grylle) on lichen-covered rock with small mackerel in its beak, UK

Steve Knell / Black Guillemot (Cepphus grylle) pair on lichen-covered rock, UK

Steve Knell / Puffin (Fratercula arctica) in flight, coming in to land. Inner Hebrides, Scotland. July

Steve Knell / Black Guillemot (Cepphus grylle) on rock with lichens and sea thrift, UK

Steve Knell / Black Guillemot (Cepphus grylle) on rock, calling. UK