Ben Hall / Marine iguana (Amblyrhynchus cristatus) on rock taken with slow shutter speed to show motion in the water, Galapagos islands, May.

Wim van den Heever / Nile crocodile (Crocodylus niloticus) still in the fast flowing water of the Mara River, taken with slow shutter speed. Masai Mara, Kenya.

Rob Jordan / 2020VISION / Arctic tern (Sterna paradisaea) in flight at twilight using slow shutter speed to accentuate blur and motion in low-key light, Inner Farne, Farne Islands, Northumberland, July.

Karine Aigner / Slow shutter portrait of an African wild dog (Lycaon pictus). Malilangwe Wildlife Reserve, Zimbabwe.

Andy Trowbridge / Looking up Otira Valley with Otira River in the foreground. Slow shutter speed showing movement of the water. Arthur's Pass National Park, Southern Alps, New Zealand. May.

SCOTLAND: The Big Picture / Common toad (Bufo bufo) on fallen tree over river, Atlantic rainforest, Glen Nant, Scotland, UK, September.

Doug Perrine / Hot lava from the 61G flow from Kilauea Volcano enters the ocean from the open end of a lava tube at the Kamokuna entry in Hawaii Volcanoes National Park, producing a great cloud of steam; a slow came...

Pal Hermansen / Red necked phalarope (Phalaropus lobatus) preening, taken with slow shutter speed, Iceland, June.

Wim van den Heever / Giant anteater (Myrmecophaga tridactyla) walking through an open field, taken with slow shutter speed. Pantanal, Brazil.

Wim van den Heever / Japenese cranes (Grus japonensis) in courtship display, taken with slow shutter speed, Hokkaido Japan. March.

Ben Hall / The Strid, River Wharfe, slow shutter speed showing movement of the water, Bolton Abbey Estate, Wharfedale, North Yorkshire, August 2015

Ben Hall / The Strid, River Wharfe, slow shutter speed showing movement of the water, Bolton Abbey Estate, Wharfedale, North Yorkshire, August 2015