Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Christophe Courteau / Blue footed booby displaying {Sula nebouxii} Galapagos.

Ross Hoddinott / Robin (Erithacus rubecula) displaying in snow, nr Bradworthy, Devon, UK. December 2010, HIGHLY COMMENDED, ANIMAL PORTRAITS, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR COMPETITION

Wild Wonders of Europe / E. Haarberg / Male Black grouse (Tetrao tertrix) displaying, expiring warm air on a cold morning, Bergslagen, Sweden, April 2009

Tony Wu / Sperm whale (Physeter macrocephalus), ten swimming together in gathering of more than 50 individuals. Whales with flatulence and defecating. Sri Lanka.

Ann & Steve Toon / Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August

Nick Garbutt / Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January.

Dave Bevan / Common wasp (Vespula vulgaris) adding material to nest, Carmarthenshire, Wales, UK, July.

Cyril Ruoso / Long-tailed macaque (Macaca fascicularis) grooming, Koram island Khao Sam Roi Yot National Park, Thailand.

Frederique Olivier / Close-up of a pair of Black browed albatrosses (Thalassarche melanophris) mutual grooming, New Island, Falkland Islands.

Dietmar Nill / European hornet (Vespa crabro) fanning nest with wings to regulate the temperature, Germany, September.

Konrad Wothe / Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.