Thomas Marent / Mindanao Splash Frog (Staurois natator) raising foot during display, Bako National Park, Sarawak, Borneo, Malaysia, digital composite

Chien Lee / Sabah Splash Frog (Staurois latopalmatus), Sarawak, Borneo, Malaysia

Nick Garbutt / Black-spotted Rock Frog (Staurois natator). Centre of Maliau Basin, Sabah's 'Lost World', Borneo.

Jouan & Rius / Black spotted rock frog (Staurois natator) Gunung Gading NP, Borneo, Sarawak, Malaysia

Chien Lee / Mindanao Splash Frog (Staurois natator), Danum Valley Conservation Area, Sabah, Borneo, Malaysia

Chien Lee / Mindanao Splash Frog (Staurois natator), Mulu National Park, Borneo, Malaysia

Chien Lee / Borneo Splash Frog (Staurois tuberilinguis), Mount Kinabalu National Park, Borneo, Malaysia

Chien Lee / Sabah Splash Frog (Staurois latopalmatus), Gunung Mulu National Park, Sarawak, Malaysia

Chien Lee / Sabah Splash Frog (Staurois latopalmatus), Gunung Mulu National Park, Sarawak, Malaysia

Doug Wechsler / Green spotted rock frog {Staurois tuberilinguis} Montane forest, Borneo

Nick Garbutt / Green-spotted rock frog (Staurois tuberilinguis) on stream side rock. Danum Valley, Sabah, Borneo, Malaysia

Thomas Marent / Mindanao Splash Frog (Staurois natator) camouflaged against mossy rocks, Danum Valley Conservation Area, Malaysia

Thomas Marent / Mindanao Splash Frog (Staurois natator) on mossy rocks, Danum Valley Conservation Area, Malaysia

Thomas Marent / Mindanao Splash Frog (Staurois natator), Danum Valley Conservation Area, Malaysia

Tony Heald / Black-spotted Rock Frog [Staurois natator] Danum Valley, Sabah, Borneo, September

Doug Wechsler / Rock skipper {Staurois latopalmatus} on rock in river rapids, W Kalimantan, Indonesia

Doug Wechsler / Rock skipper sitting on rock in river rapids {Staurois latopalmatus} West Kalimantan, Borneo, Indonesia