John Chan / Killdeer (Charadrius vociferus) stamping its feet to create vibrations in the sand as it forages for insects, Southern California, USA, August.

Guy Edwardes / Allen's / Lesser gallinule (Porphyrio alleni) walking on waterlilies, Chobe River, Botswana.

George Sanker / Purple gallinule (Porphyrio martinica) moving across waterlily covered surface, Everglades National Park, Florida, USA, March

Ross Hoddinott / Tarr Steps across the River Barle in autumn, nr Withypool, Exmoor NP, Somerset, UK.

Laurie Hedges / Slow motion clip of an adult female Baird's tapir (Tapirus Bairdii) walking through a rainforest, stepping over a tree root, Corcovado National Park, Costa Rica.

Ingo Arndt / Mountain Lion (Puma concolor) stepping into Guanaco (Lama guanicoe) tracks in snow during stalk, Torres del Paine National Park, Patagonia, Chile

Neil Aldridge / Red fox (Vulpes vulpes) juvenile showing signs of mange in its coat and eyes, stepping out from its den behind a garden shed, Bristol, England, UK. June 2015. Winner of the Portfolio Category of Natur...

David Pattyn / Grey Heron (Ardea cinerea) standing at the edge of a pond while preening its feathers. Valkenhorst nature reserve, Valkenswaard, The Netherlands June

Philippe Clement / Grey wolf (Canis lupus) pack stepping over a fallen tree trunk in the snow, Bavarian Forest National Park, Germany, March. Captive.

Nick Upton / Greylag goose (Anser anser) stepping through ice whilst emerging from a partly frozen marshland pool, Gloucestershire, England, UK, January.

Nick Upton / Eurasian beaver (Castor fiber) stepping over a dam to reach a beaver pond in a large woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, June. Taken by a remote...

Gabriel Rojo / White-banded mockingbird, (Mimus triurus) walking, Calden Forest, La Pampa , Argentina