Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

Dave Watts / Wedge-tailed eagle {Aquila audax fleayi} tasmanian sub species, adult portrait, Australia

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) fourteen year old sub-adult male, named Fundi, making submissive facial gesture, Gombe National Park, Tanzania

Nick Garbutt / Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December.

Tony Wu / Bowhead whale (Balaena mysticetus), group socialising in shallow water, aerial view. Part of larger group of more than 50 whales congregating, Sea of Okhotsk sub-population, Russia.

Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.

Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.

Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.

SCOTLAND: The Big Picture / Mountain hare (Lepus timidus), sub-adult in summer coat on moorland, Scotland, UK, August.

Franco Banfi / Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico.

Franco Banfi / California sealion (Zalophus californianus) swimming. Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico

Franco Banfi / Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.