Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) fourteen year old sub-adult male, named Fundi, making submissive facial gesture, Gombe National Park, Tanzania

Ann & Steve Toon / Spotted eagle owl (Bubo africanus) subadult at nigh by water, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Alan Murphy/ BIA / Northern Caracara (Caracara cheriway) chasing sub-adult with prey, Texas

Tui De Roy / Short tailed albatross (Phoebastria albatrus) subadult with fish-hook and monofilament line embedded in throat. Tsubamezaki, Torishima Island, Japan. December.

Ingo Arndt / Baya weaver (Ploceus philippinus) subadult birds practising building 'play nest', Singapore.

Ingo Arndt / Desert locust subadult {Schistocerca gregaria} Africa

Ingo Arndt / Gelada baboon subadult males grooming {Theropithecus gelada} Simien Mt NP, Ethiopia

Yva Momatiuk & John Eastcott / Ostrich (Struthio camelus) sub-adults at waterhole in dry season, Etosha National Park, Namibia

Gerrit Vyn / Subadult male Wild Turkey (Meleagris gallopavo) in alternate (breeding) plumage. Hidalgo County, Texas. March.

Gerry Ellis / Orangutan (Pongo pygmaeus) sub-adult male in trees in haze caused by forest clearing, West Kalimantan, Borneo, Indonesia. October, 2015

Greg Oakley / Australian King Parrot (Alisterus scapularis) sub-adult male, Victoria, Australia

Graeme Guy / White-backed Vulture (Gyps africanus) and sub-adult, Masai Mara, Kenya