Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) fourteen year old sub-adult male, named Fundi, making submissive facial gesture, Gombe National Park, Tanzania

Martin Willis / Budgerigar (Melopsittacus undulatus) father with begging chick at nest cavity, Queensland, Australia

Willi Rolfes/ BIA / Alpine Marmot (Marmota marmota) parent feeding with begging young, Hohe Tauern National Park, Austria

Heike Odermatt / King Penguin (Aptenodytes patagonicus) chick begging for food, Saunders Island, Falkland Islands

Edwin Giesbers / Barbary macaque (Macaca sylvanus) baring teeth as a sign of submission, Gibraltar Nature Reserve, Gibraltar,

Suzi Eszterhas / Snowy Egret (Egretta thula) four week old chick begging for food, Sonoma County, California

Bruno de Faveri / Arctic Tern (Sterna paradisaea) with begging chick, Farne Islands, England, United Kingdom

Michael Quinton / Northern Flicker (Colaptes auratus) entering nest cavity with begging chicks, Alaska

Michael Quinton / Northern Flicker (Colaptes auratus) at nest cavity with begging chicks, Alaska

Roland Seitre / Eastern Wolf (Canis lupus) showing submissive behavior to dominant individual, Omega Park, Montebello, Quebec, Canada

Laurent Geslin / Black backed jackal, submission {Canis mesomelas} Skeleton Coast, Namibia

Laurent Geslin / Two male urban Red foxes, mouths open, one in submissive posture {Vulpes vulpes} London, UK