Sandra Bartocha / Sun rays shining through European beech woodland (Fagus sylvatica). Serrahn, Muritz-National Park, World Natural Heritage site, Germany, Europe. November 2015.
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Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.
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Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) with young chicks on back, Cardiff, UK, April.
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Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
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Floris van Breugel / Autumnal landscape with snow, Arrigetch Peaks region of Brooks Range, Gates of the Arctic National Park, Alaska, USA, August 2014.
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Floris van Breugel / Black-billed shrike-tyrant (Agriornis montanus) in the snow with Mount Fitz Roy visible beyond. Patagonia, Argentina, June 2014.
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Floris van Breugel / Salto Grande waterfall with Paine Grande beyond, Torres del Paine National Park, Chile, June 2014.
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Ross Hoddinott / Basalt columns at Vik, the southernmost village in Iceland. February 2014.
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Anup Shah / Spotted hyena (Crocuta crocuta) preparing to carry a pup aged 2-3 months. Masai Mara National Reserve, Kenya, July
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Anup Shah / Lion (Panthera leo) male nuzzling lioness, Masai Mara National Reserve, Kenya, July
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Danny Green / Grey Squirrel (Sciurus carolinensis) sunlit portrait. Leicestershire, UK, November.
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Erlend Haarberg / Sunlit ridges against volcanic scree beneath a low cloud. Eyjafjallajokull, Iceland, April 2010. Highly commended in Wildlife Photographer of the Year 2011 competition, Wild Places category.