Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February

Richard Du Toit / Burchell's Zebra (Equus burchellii) pair, Rietvlei Nature Reserve, Gauteng, South Africa

Gavin Hellier / Two buddhist monks relaxing in the Bayon Temple, Angkor Wat, Siem Reap, Cambodia 2010. Model released.

Nick Turner / Women and children having a picnic high up on Selsley Common unimproved grassland, Stroud, Gloucestershire, UK, August 2011. Model released

Nick Turner / A couple relaxing amongst bluebells (Hyacinthoides non-scripta) in springtime woodland, Upper Soudley, Forest of Dean, Gloucestershire, May.

Jeff Wilson / Director, Chadden Hunter, standing on wind eroded rock talking on radio to base camp, Dry Valleys, Antarctica, December 2009. Taken on location for the BBC series, Frozen Planet.

Matthew Maran / Hikers talking as they warm themselves by a campfire on a beach. The West Coast Trail, Pacific Rim National Park, Vancouver Island, Canada, September 2010. Models released.

Yva Momatiuk & John Eastcott / Grizzly Bear (Ursus arctos horribilis) pair of young males fighting, Katmai National Park, Alaska

Olga Kamenskaya / View through door of people talking, whilst on diving expedition. Upper reaches of the Lena River Baikalo-Lensky Reserve, Siberia, Russia. August 2018.

Staffan Widstrand / Wild Wonders of China / Hill myna (Gracula religiosa), caught in the wild, for sale at a truckstop, Shaanxi, China.

Cyril Ruoso / Shoebill (Balaeniceps rex) biologist, Elijah Mofya, on radio in tree, Bangweulu Wetlands, Zambia

Jabruson / Ouled Rachid tribeswoman talking, Kashkasha village near Zakouma National Park, Chad, 2010.