Giri Cavale / Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
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Vincent Grafhorst / Hippopotamus (Hippopotamus amphibius) bull marking territory by flicking dung, Chobe National Park, Botswana
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Wild Wonders of Europe / E Haarberg / Chamois (Rupicapra rupicapra) male marking his territory in rutting season, Gran Paradiso National Park, Italy, November 2008
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Anup Shah / Simien jackal / Ethiopian wolf {Canis simensis} marking territory by scraping, Bale Mountains, Bale NP, Ethiopia
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Ann & Steve Toon / Cheetah (Acinonyx jubatus) spraying / scentmarking tree, Zimanga private game reserve, Kwaqulu-Natal, South Africa.
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Sandesh Kadur / Snow leopard (Unica unica) scent marking against boulder in rocky landscape in Himalayas. Ule, Ladakh, India. March 2018. Sequence 2/2.
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Yashpal Rathore / Bengal Tiger (Panthera tigris), Spray marking on tree to establish its territory, Jim Corbett National Park, India
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Tony Heald / Lion (Panthera leo) male, marking territory, Etosha National Park Namibia.
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Oriol Alamany / Snow leopard (Panthera uncia) male at dusk marking territory on a rocky slope at 4000 m in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India
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Giri Cavale / Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
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Giri Cavale / Melanistic leopard / Black panther (Panthera pardus) male in tree, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
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Giri Cavale / Leopard (Panthera pardus) male chasing away a melanistic leopard / black panther, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
