John Chan / American avocet (Recurvirostra americana) foraging in salt marsh, Southern California, USA.
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John Shaw / Brown bear (Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August.
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Pedro Narra / Aerial view of low tide in Sado estuary, Portugal. November
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Guy Edwardes / Gatklettur, Arnarstapi, Snaefellsnes Peninsula, Iceland, March 2017.
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Guy Edwardes / Rock formations at Kimmeridge Bay, Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
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Guy Edwardes / Luskentyre beach with three people wading in water, Isle of Harris, Outer Hebrides, Scotland, UK. June 2012.
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Guy Edwardes / Saltmarsh on the Isle of Harris, Outer Hebrides, Scotland, UK. November 2016.
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Guy Edwardes / Saltmarsh. Isle of Harris, Scotland, UK. March, 2015.
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Dave Bevan / Grey plover (Pluvialis squatarola) eating a marine flatworm, Norfolk, England, UK, February.
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Nick Upton / Common starfish (Asterias rubens) moving in a rockpool, real time, Cornwall, England, UK, September.
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Nick Upton / Common prawns (Palaemon serratus) and a Rock goby (Gobius paganellus) feeding on a dead Montagu's crab (Xantho hydrophilus) in a rockpool, Cornwall, England, UK, September.
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Nick Upton / Rock goby (Gobius paganellus) stealing food from a scavenging Common prawn (Palaemon serratus) in a rockpool, Cornwall, England, UK, September.