John Shaw / Brown bear (Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August.
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) digging for clams on tidal flats, Lake Clark National Park, Alaska.
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Ernie Janes / Low tide on the Wash, Snettisham, Norfolk, England, UK, February.
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) cub on mothers back walking on tidal flats, Lake Clark National Park, Alaska
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David Tipling / 2020VISION / Bar-tailed Godwit (Limosa lapponica) foraging for tidal-flat worm. Norfolk, January. Sequence 2 of 2.
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David Tipling / 2020VISION / Bar-tailed Godwit (Limosa lapponica) foraging for tidal-flat worm. Norfolk, January. Sequence 1 of 2.
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John Chan / Green heron (Butorides virescens) stalking and catching a Topsmelt silverside (Atherinops affinis) on the edge of a tidal flat, Bolsa Chica Ecological Reserve, California, USA, September.
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Nick Upton / Common prawns (Palaemon serratus) scavenging on a damaged Flat top shell (Gibbula umbilicalis) in a rockpool, Cornwall, England, UK, September.
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Nick Upton / Cushion star (Asterina gibbosa) moving in a rockpool near some Flat top shells (Gibbula umbilicalis), Cornwall, England, UK, September.
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Ingo Arndt / Grizzly Bear / Coastal Brown Bear (Ursus arctos horribilis) mother with spring cub digging for clams on tidal flats, Lake Clark National Park, Alaska, USA. June.
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Ingo Arndt / Grizzly Bear / Coastal Brown Bear (Ursus arctos horribilis) mother with spring cubs digging for clams on tidal flats, Lake Clark National Park, Alaska, USA. June.
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Ingo Arndt / Grizzly Bear / Coastal Brown Bear (Ursus arctos horribilis) mother with spring cubs digging for clams on tidal flats, Lake Clark National Park, Alaska, USA. June.