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Waves breaking against sea wall and cliffs, Seaford, Sussex, England, UK, 15th February 2014.
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Mark Taylor / Waves breaking against sea wall and cliffs, Seaford, Sussex, England, UK, 15th February 2014.
Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
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Mark Hamblin / 2020VISION / Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
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Mark Hamblin / 2020VISION / Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
Shrimp fishermen's trailers inundated by flood water during an exceptionally high tide near Cark in Cartmell on Morecambe Bay, Cumbria, England, UK. February 2014
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Ashley Cooper / Shrimp fishermen's trailers inundated by flood water during an exceptionally high tide near Cark in Cartmell on Morecambe Bay, Cumbria, England, UK. February 2014
Land crab (Gecarcinidae) pausing just above high tide mark, Savusavu, Fiji.
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David Fleetham / Land crab (Gecarcinidae) pausing just above high tide mark, Savusavu, Fiji.
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013.
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SCOTLAND: The Big Picture / Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013.
Tyre tracks in sand on beach as low tide, Crotoy, Somme Bay, Pas de Calais, France, July 2015.
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Pascal Tordeux / Tyre tracks in sand on beach as low tide, Crotoy, Somme Bay, Pas de Calais, France, July 2015.
Tidal surge damage at Cley Beach and Nature Reserve, Norfolk, December 2013.
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Ernie Janes / Tidal surge damage at Cley Beach and Nature Reserve, Norfolk, December 2013.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) on beach feeding on cockles cracked opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) on beach feeding on cockles cracked opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468311
Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468283
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468281
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) on beach using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468279
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468278
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468276
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) feeding on cockles opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468275
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) feeding on cockles opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) feeding on cockles opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468274
Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) feeding on cockles opened using stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468258
Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open cockles at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468230
Mark MacEwen / Burmese long tailed macaques (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
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Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468193
Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468192
Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468191
Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468190
Mark MacEwen / Male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) feeding on cockles opened with stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468165
Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) feeding on cockles opened with stone tools at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468132
Mark MacEwen / Burmese long tailed macaque (Macaca fascicularis aurea) using stone tools to open Oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
01468124
Mark MacEwen / Adult male Burmese long tailed macaque (Macaca fascicularis aurea) using stone tool to open oysters at low tide, Kho Ram, Khao Sam Roi Yot National Park, Thailand.
Heron Island Reef Wall at low tide with exposed corals, southern Great Barrier Reef, Queensland, Australia.
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Mark MacEwen / Heron Island Reef Wall at low tide with exposed corals, southern Great Barrier Reef, Queensland, Australia.
Heron Island Reef Wall at low tide, exposing corals southern Great Barrier Reef, Queensland, Australia.
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Mark MacEwen / Heron Island Reef Wall at low tide, exposing corals southern Great Barrier Reef, Queensland, Australia.
Staghorn Coral (Acropora cervicornis) at low tide, Great barrier reef, Heron Island, Australia.
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Mark MacEwen / Staghorn Coral (Acropora cervicornis) at low tide, Great barrier reef, Heron Island, Australia.
Exposed Staghorn coral (Acropora cervicornis) at low spring tide, Heron Island, southern Great Barrier Reef, Queensland, Australia.
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Mark MacEwen / Exposed Staghorn coral (Acropora cervicornis) at low spring tide, Heron Island, southern Great Barrier Reef, Queensland, Australia.
Seaweed washed up on tide line of beach. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, March.
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Mark Hamblin / 2020VISION / Seaweed washed up on tide line of beach. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, March.
Seaweed washed up on tide line of beach. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, March.
01393973
Mark Hamblin / 2020VISION / Seaweed washed up on tide line of beach. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, March.
Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
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Mark Hamblin / 2020VISION / Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
01393969
Mark Hamblin / 2020VISION / Close-up of shells on tide line. Sandyhills Bay, Solway Firth, Dumfries and Galloway, Scotland, UK, February 2012.
Beach Stone-curlew (Esacus magnirostris) egg in nesting depression above the high tide mark, Toolakea Beach, Queensland, Australia
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Martin Willis / Beach Stone-curlew (Esacus magnirostris) egg in nesting depression above the high tide mark, Toolakea Beach, Queensland, Australia
East coast of Lundy Island at low tide, UK.
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Mark Taylor / East coast of Lundy Island at low tide, UK.
Pantropical spotted dolphin {Stenella attenuata} Gulf of Mexico Atlantic
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Todd Pusser / Pantropical spotted dolphin {Stenella attenuata} Gulf of Mexico Atlantic
Military River Frog (Thoropa miliaris) the only frog in the world to hunt in tide-pools, Atlantic Forest, near Sao Paulo, Brazil
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Mark Moffett / Military River Frog (Thoropa miliaris) the only frog in the world to hunt in tide-pools, Atlantic Forest, near Sao Paulo, Brazil
Military River Frog (Thoropa miliaris) the only frog in the world to hunt in tide-pools, observed by biologist Richardo Sowaya Atlantic Forest, near Sao Paulo, Brazil
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Mark Moffett / Military River Frog (Thoropa miliaris) the only frog in the world to hunt in tide-pools, observed by biologist Richardo Sowaya Atlantic Forest, near Sao Paulo, Brazil
High tide mark shows oil on rocky shore spilled by Exxon Valdez oil spill, Prince William Sound, Alaska
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Flip Nicklin / High tide mark shows oil on rocky shore spilled by Exxon Valdez oil spill, Prince William Sound, Alaska
Dead seagrass at high tide mark on beach, Western Australia
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Elizabeth White / Dead seagrass at high tide mark on beach, Western Australia
Cuttlefish bones on high tide mark {Sepioidea} Coral Bay, Western Australia
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Elizabeth White / Cuttlefish bones on high tide mark {Sepioidea} Coral Bay, Western Australia
Adult Eurasian otter (Lutra lutra) defecating on shoreline to mark territory before walking away with her pup following close behind, Norway.
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Ismaele Tortella / Adult Eurasian otter (Lutra lutra) defecating on shoreline to mark territory before walking away with her pup following close behind, Norway.