Alex Mustard / Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...
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Jack Dykinga / Burrowing owl (Athene cunicularia) fledgling aged one month tilting head to judge distance. Marana, Arizona, USA. May.
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Andy Rouse / Golden oriole (Oriolus oriolus) pair, male perched on stump, female flying above. Ranthambhore National Park, India.
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Laurie Hedges / Drone shot tracking over oil palm plantations and rainforest edge, showing areas of deforestation, Rondonia, Brazil, 2019.
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Derek Galon / Drone shot tilting down to Victoria Falls, water is milky blue as it contains sulphur from Boiling Lake, Dominica, 2019.
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Ashley Cooper / Second World War lookout post leaning over eroding cliff near Aldbrough, Yorkshire, England, UK. August 2013.
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Ashley Cooper / Second World War pill box on the beach, this was originally on the cliffs above but fell due to coastline erosion. Aldbrough, Yorkshire, England, UK. August 2013.
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Ashley Cooper / Second World War lookout post leaning over eroding cliff near Aldbrough, Yorkshire, England, UK. August 2013.
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Philippe Clement / Tilting shot of Beech trees (Fagus sylvatica) in autumn, Bavarian Forest National Park, Germany, October
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John Waters / Shoal of Dace (Leuciscus leuciscus) feeding in the River Kennet, Berkshire, England, UK, March.
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Frederique Olivier / Tilting tracking shot of a mountain and its reflection, seen from Gerlache Strait, Antarctic Peninsula.
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Frederique Olivier / Tilting shot upwards from a reflection of Adelie penguins (Pygoscelis adeliae) in water to a group resting on sea ice, Antarctica.
