Ingo Arndt / Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December

Ingo Arndt / Tisza mayfly (Palingania longicauda) hanging on branch and moulting, River Tisza, Hungary, June

Ingo Arndt / Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December

Ingo Arndt / Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September

John Waters / Pair of Brown trout (Salmo trutta) spawning, with others eating eggs, River Kennet, Berkshire, England, UK. March.

John Waters / Female Grayling (Thymallus thymallus) pursued by several males, River Kennet, Berkshire, England, UK. March.

John Waters / Pair of Brown trout (Salmo trutta) spawning, with another male trying to fertilise eggs River Kennet, Berkshire, England, UK. February.

John Waters / Brook lamprey (Lampetra planeri) looking for food, River Kennet, Berkshire, England, UK. January.

Francisco Marquez / Griffon vultures (Gyps fulvus) and European black vultures (Aegypius monachus) feeding on a dead deer, Cabaneros National Park, Montes de Toledo, Spain, May.

Francisco Marquez / Juvenile Lammergeier (Gypaetus barbatus) feeding on a bone in snow, Ordesa y Monte Perdido National Park, Spain, April.

Mark Hamblin / 2020VISION / Two male Black grouse (Tetrao tetrix) fighting at lek site, with mountain backdrop, Cairngorms NP, Scotland, UK, April 2010

Mark Hamblin / 2020VISION / Red squirrel (Sciurus vulgaris) adult foraging around a woodland pool, Cairngorms NP, Scotland, UK, November 2011