Ingo Arndt / Female Puma (Puma concolor) with two cubs aged around four months, Torres del Paine National Park, Patagonia, Chile, July.

Ingo Arndt / Close-up of a Guanaco (Lama guanicoe) feeding, Torres del Paine National Park, Patagonia, Chile, December.

Ingo Arndt / Aerial shot over a herd of Guanacos (Lama guanicoe), Torres del Paine National Park, Patagonia, Chile, April.

Terry Whittaker / Man collecting seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down cliff to collect seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being raised up cliff to collect seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man sorting seabird eggs, including those of Common guillemots (Uria aalge) into egg boxes, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.

John Chan / Great horned owl (Bubo virginianus) chick watching for parents to return with food, Southern California, USA, June.

John Chan / Peregrine falcon (Falco peregrinus) plucking Dowitcher (Limnodromus) prey, Southern California, USA.

John Chan / American avocet (Recurvirostra americana) foraging in salt marsh, Southern California, USA.

John Chan / Killdeer (Charadrius vociferus) stamping its feet to create vibrations in the sand as it forages for insects, Southern California, USA, August.

John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019

John Chan / Flock of Western sandpipers (Calidris mauri) taking flight from salt marsh, Southern California, USA.

John Chan / California sea lion (Zalophus californianus) feeding on a Striped bass (Morone saxatilis), Southern California, USA.

Morley Read / Column of Leaf cutter ants (Atta) carrying cut pieces of leaves along the forest floor towards therr nest, Orellana Province, Ecuador, 2018. (non-ex)

Morley Read / Leaf frog (Agalychnis hulli) blinking its eyes and turning around, Orellana Province, Ecuador. (non-ex)

Morley Read / Developing embryo of the Ecuadorian monkey frog (Phyllomedusa ecuatoriana), showing heart beating and blood flowing, Morona Santiago Province, Ecuador, (non-ex)

Morley Read / Close-up of an Amazon leaf frog (Cruziohyla Craspedopus) tadpole developing in egg, Ecuador. (non-ex)

Sandesh Kadur / Slow motion clip of Amur falcons (Falco amurensis) congregating at roost site during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Espen Bergersen / Aerial shot of a fishing boat catching Atlantic herring (Clupea harenguis), with Killer whales (Orcinus orca) and Humpback whales (Megaptera novaeangliae) swimming around the boat and large numbers of...

Espen Bergersen / Aerial shot of a fishing boat catching Atlantic herring (Clupea harenguis), with Killer whales (Orcinus orca) and Humpback whales (Megaptera novaeangliae) swimming around the boats and large numbers o...

Espen Bergersen / Aerial shot of fishing boats catching Atlantic herring (Clupea harenguis), with Killer whales (Orcinus orca) swimming around the boats, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a shoal of Atlantic herring (Clupea harenguis), hunting, with one Killer whale tail slapping to stun the Herring, Sommaroy, Troms, Norwa...

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming into a huge shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming near to a huge shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a large shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of a huge shoal of Atlantic herring (Clupea harenguis), with a Humpback whale (Megaptera novaeangliae) nearby, Sommaroy, Troms. Norway, January.

Espen Bergersen / Aerial shot of a Killer whale (Orcinus orca) spyhopping several times close to land, with people watching and taking pictures, Sommaroy, Troms. Norway, January.

Espen Bergersen / Aerial shot of three Killer whales (Orcinus orca) swimming close to the surface, Troms, Norway, August.

Espen Bergersen / Aerial shot of Killer whales (Orcinus orca) near the surface, showing mating behaviour including a male Killer whale displaying penis, Troms, Norway, August.

Espen Bergersen / Aerial shot of Killer whales (Orcinus orca) near the surface, showing mating behaviour, Troms, Norway, August.

Espen Bergersen / Aerial shot of a fishing boat with net full of Atlantic herring (Clupea harengus), with Killer whales (Orcinus orca) swimming around the outside, Skjervoy, Troms, Norway, November.

Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) hunting Atlantic herring (Clupea harengus), Troms, Norway, October.

Espen Bergersen / Aerial shot of a pod of Killer whales (Orcinus orca) swimming near the surface, Troms, Norway, October.

Espen Bergersen / Aerial shot of five Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of a group of Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, one whale creating a bubble n...

Espen Bergersen / Aerial shot of five Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus), Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of three Humpback whales (Megaptera novaeangliae) swimming into a shoal of Atlantic herring (Clupea harengus), preparing to lunge feed, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) swimming near to a shoal of Atlantic herring (Clupea harengus), Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of a Humpback whale (Megaptera novaeangliae) swimming showing belly, Troms, Norway, January.

Espen Bergersen / Aerial shot of a Humpback whale (Megaptera novaeangliae) showing belly and blowing, Troms, Norway, January.

Espen Bergersen / Aerial shot of a Humpback whale (Megaptera novaeangliae) diving and showing tail fluke, Troms, Norway, January.

Espen Bergersen / Aerial shot of a large group of Humpback whales (Megaptera novaeangliae), Troms, Norway, January.

Espen Bergersen / Aerial shot of a large group of Humpback whales (Megaptera novaeangliae), Troms, Norway, January.

Espen Bergersen / Aerial shot of two Humpback whales (Megaptera novaeangliae) coming to the surface to breathe, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of a group of Killer whales (Orcinus orca) tail slapping, shot pans to the right, Skjervoy, Troms, Norway, November.

Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) carousel feeding close to Tromvik harbour, Troms, Norway, December.

Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) hunting Atlantic herring (Clupea harengus) using carousel feeding, with herring jumping out of the water and gulls (Larus) circling above, Tromvik, Troms, No...

Espen Bergersen / Aerial shot of a female Humpback whale (Megaptera novaeangliae) and calf surrounded by Killer whales (Orcinus orca), mother hits sideways with her tail fluke to drive Killer whales away, Skjervoy, Tro...

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) spyhopping, with a snowy landscape in the background, Skjervoy, Troms, Norway, December.

Espen Bergersen / Aerial shot of a small pod of Killer whales (Orcinus orca) at surface, with snowy landscape in the background, Skjervoy, Troms, Norway, December.

Espen Bergersen / Aerial shot of a pod of Killer whales (Orcinus orca) carousel feeding, Skjervoy, Troms, Norway, November.

Espen Bergersen / Aerial shot of a pod of Killer whales (Orcinus orca) carousel feeding, drone moving closer, Skjervoy, Troms, Norway, November.

John Waters / Large murmuration of Common starlings (Sturnus vulgaris) swirling over reedbed roost at sunset, Ham Wall RSPB Reserve, Somerset, England, UK, January.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

David Perpinan / Tracking shot of a male Pheasant (Phasianus colchicus) displaying to an unreceptive female, Barcelona, Spain, May.

David Perpinan / Male Mediterranean tree frog (Hyla meridionalis) calling, showing vocal sac inflating, Barcelona, Spain, May. Filmed at night with an infrared camera.

David Perpinan / Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.

David Perpinan / Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, with another chick leaning in, chased away by adult, Barcelona, Spain, May.

David Perpinan / Grey heron (Ardea cinerea) regurgitating a fish for its chick, Barcelona, Spain, May.

Brian Bevan / Juvenile Green woodpecker (Picus viridis) leaving nest hole, shot looking up trunk, Bedfordshire, England, UK, June.

Brian Bevan / Male Green woodpecker (Picus viridis) feeding young at nest hole, shot looking up trunk, Bedfordshire, England, UK, June.

Brian Bevan / Pantaloon bee (Dasypoda hurtles) entering burrow with pollen, Bedfordshire, England, UK, August.

Brian Bevan / Long tailed tit (Aegithalos caudates) nest building, Bedfordshire, England, UK, April.

Brian Bevan / Slow motion clip of a male Kingfisher (Alcedo atthis) taking fish into nest hole before exiting backwards, Bedfordshire, England, UK, July.

Brian Bevan / Brown hare (Lepus europeans) feeding on barley, Bedfordshire, England, UK, June.

John Cancalosi / American toads (Anaxyrus americanus), several males competing to mate with female, Maryland, USA, April.

Kerstin Hinze / Red crossbill (Loxia curvirostra) feeding from a spruce cone, Bavarian Forest National Park, Germany, October.

Kim Taylor / Slow motion clip of a male Common blackbird (Turdus merula) bathing in a birdbath, Surrey, England, UK, October.

Solvin Zankl / Common mole (Talpa europaea) feeding on a grub underground, Germany. Live, captive animal.

Kerstin Hinze / Two Black-bellied hamsters (Cricetus cricetus) fighting, Vienna, Austria, October.

Kerstin Hinze / Male Common european toad (Bufo bufo) in amplexus with a dead fish, Lower Saxony, Germany, April.

Kerstin Hinze / Painted lady (Vanessa cardui) nectaring, Bavarian Forest National Park, Germany, June.

Kerstin Hinze / Male Capercaillie (Tetrao urogallus) displaying, Berchtesgaden National Park, Bavarian Alps, Germany, April.

Kerstin Hinze / Male Capercaillie (Tetrao urogallus) displaying, Berchtesgaden National Park, Bavarian Alps, Germany, April.

Kerstin Hinze / Dipper (Cinclus cinclus) foraging in a stream, Harz National Park, Saxony-Anhalt, Germany, March.

Remi Masson / Pike (Esox lucius) hunting in a shoal of Perch (Perca fluviatilis), Lake Neuchatel, Switzerland. June.

Remi Masson / Large Pike (Esox lucius) more than a meter long, resting on a lake bed, Lake Neuchatel, Switzerland. June.

Philippe Clement / Pair of Roe deer (Capreolus capreolus) resting and grazing in forest in summer, France, September.

Philippe Clement / Common wasp (Vespula vulgaris) feeding on jam inside a jar, Belgium, August.

Philippe Clement / Common wasp (Vespula vulgaris) feeding on jam inside a jar, Belgium, August.