Staffan Widstrand / Nyala (Tragelaphus angasi) feeding on bush, iMfolozi National Park, South Africa

Ann & Steve Toon / Nyala (Tragelaphus angasii), bulls lateral presentation display, Mkhuze Game Reserve, South Africa

Ann & Steve Toon / Nyala (Tragelaphus angasii) territorial bull scent-marking, Mkhuze game reserve, South Africa, May.

Ann & Steve Toon / Nyala (Tragelaphus angasii) with intersex syndrome, showing female coat pattern, male horns and body size, Mkhuze Game Reserve, South Africa, May.

Ann & Steve Toon / Nyala (Tragelaphus angasii) bull dominance display, Mkhuze Game Reserve, South Africa, May.

Richard Du Toit / Nyala ram (Tragelaphus angasi) lateral display, Zululand, South Africa, October

David Noton / Female Nyala (Tragelaphus angasi) in the bush, Kruger National Park, South Africa

David Noton / Female Nyala (Tragelaphus angasi) in the bush, Kruger National Park, South Africa

David Noton / Female Nyala (Tragelaphus angasi) feeding in the bush, Kruger National Park, South Africa

David Noton / Female Nyala (Tragelaphus angasi) grazing in the bush, Kruger National Park, South Africa

David Noton / Young male Nyala (Tragelaphus angasi) grazing in the bush, Kruger National Park, South Africa

David Noton / Nyala {Tragelaphus angasi} female feeding on vegetation, Kruger National Park, South africa

David Noton / Nyala bull {Tragelaphus angasi} Kruger National Park, South Africa

David Noton / Nyala bull {Tragelaphus angasi} Kruger National Park, South Africa

Philippe Clement / Nyala {Tragelaphus angasi} male, Kruger NP South Africa

Richard Du Toit / Male Nyala {Tragelaphus angasi} Phinda Resource Reserve, South Africa

Hanne & Jens Eriksen / Nyala {Tragelaphus angasi} family drinking at dried up waterhole, Mkuze GR, South Africa

Dave Watts / Nyala male drinking {Tragelaphus angasi} Mkhuzi GR, South Africa

Dave Watts / Nyala female {Tragelaphus angasi} Mkhuzi GR, South Africa

Dave Watts / Nyala young male {Tragelaphus angasi} Mkhuzi GR, South Africa

Tony Heald / Nyala male displaying, South Africa

Philippe Clement / Nyala male {Tragelaphus angasi} Kruger NP South Africa

Philippe Clement / Nyala male in woodland {Tragelaphus angasi} Kruger NP South Africa

Richard Du Toit / Nyala male, South Africa

Richard Du Toit / Nyala herd drinking at waterhole {Tragelaphus angasi} Phinda RR South Africa

Richard Du Toit / Nyala male {Tragelaphus angasi} Hluhluwe Umfolozi NP South Africa

Richard Du Toit / Two Nyala females {Tragelaphus angasi} Phinda RR South Africa

Tony Heald / Nyala male ground horning {Tragelaphus angasi} Mkuze South Africa

Tony Heald / Nyala males displaying {Tragelaphus angasi} Kruger NP South Africa