Andy Rouse / African elephant (Loxodonta africana) on plains under stormy sky, Marakele National Park, Waterberg Biosphere, South Africa

Alex Mustard / Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies.

Oscar Dewhurst / Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Yashpal Rathore / Small tree frog (Rhacophorus lateralis) at night under starry sky, Western Ghats. India. Endangered and endemic to Western Ghats.

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September

Jordi Chias / Crabeater seal (Lobodon carcinophaga) hunting under water, Antarctic Peninsula, Antarctica.

Nick Hawkins / Underneath view of a North American beaver (Castor canadensis) swimming under the ice of a frozen river. Quebec, Canada. October.

Nick Hawkins / Atlantic salmon (Salmon salar) under river ice. Quebec, Canada. October.

Juergen Freund / Moral eels, Urchins, Cuttlefish, Shrimp and other marine life living within a discarded tyre and fishing net under a ships' harbour, Maluku, Indonesia. November.

Derek Galon / Damaged forest overgrown by various vines, a typical scene in western part of Dominica, West Indies. After hurricane Maria an unprecedented number of vines took advantage of suddenly open spaces in th...

Juergen Freund / Guttering to redirect water at the Daintree Drought Experiment, where rainforest plants grown under cover to see how they respond to drought. Daintree Rainforest Observatory, northern Queensland, Aust...