Ingo Arndt / Buff tip moth (Phalera bucephala) caterpillars feeding on leaf, Hungary, June.
Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.
Jeff Vanuga / White horses of the Camargue galloping through marshes in the Camargue, France. April.
Franco Banfi / Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica.
Georgette Douwma / Big-eye fish (Priacanthus hamrur). Egypt, Red Sea.
Wim van den Heever / Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa. Runner up in the Animals in their Environment Category of the Wildl...
Sven Zacek / Roe Deer (Capreolus capreolus) herd in snowy field in Tartumaa, Estonia, March.
Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) courtship dance, Cardiff, UK, March.
Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.
David Kjaer / Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
Laurie Campbell / Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Brent Stephenson / Flock of Cape Petrels (Daption capense capense) in flight, heading away against the sky. South Shetland Islands, Antarctica, December.
Chadden Hunter / King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Martin Gabriel / Four Griffon Vultures (Gyps fulvus) in flight, silhouetted overhead. Monfrague National Park, Extremadura, Spain, March.
Kristel Richard / Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croy's estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Sandesh Kadur / Great-tailed Grackle (Quiscalus mexicanus) roosting on electric lines along Highway 77, Brownsville, Texas, USA. March
Grzegorz Lesniewski / Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November.
Grzegorz Lesniewski / Herd of Carpathian Pony / Hucul (Equus caballus) in snow. Bieszczady National Park, the Carpathians, Poland, March.
Doug Perrine / Swarm / aggregation of venomous Moon jellyfish (Aurelia aurita) Port Fidalgo, Prince William Sound, USA, July
Jurgen Freund / School of Yellow line / Yellowstripe scads (Selaroides leptolepis) Misool, Raja Ampat, West Papua, Indonesia
Jurgen Freund / Schooling Blackfin barracuda (Sphyraena qenie) West New Britain, Papua New Guinea
Jurgen Freund / School of Yellow line scads (Selaroides leptolepis) Misool, Raja Ampat, West Papua, Indonesia
Jurgen Freund / School of Yellow line scads (Selaroides leptolepis). Misool, Raja Ampat, West Papua, Indonesia
Jurgen Freund / Pod of Indo-Pacific bottlenose dolphins (Tursiops aduncus) just below sea surface in flat calm waters. Raja Ampat, West Papua, Indonesia, February 2010
Colin Varndell / Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October.
Edwin Giesbers / Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January.
Sergey Gorshkov / Salmon (Salmonidae) migrating to spawn in Lake Kuril, Kamchatka, Far East Russia, August
Orsolya Haarberg / Red deer (Cervus elaphus) herd wading through water in a flooded forest, Danube-Drava National Park, Hungary.
Solvin Zankl / Pair of Common european toads (Bufo bufo) swimming together in pond, Germany
Kim Taylor / Wood Ant (Formica rufa) workers defending the nest by ejecting droplets of formic acid, UK.
Barrie Britton / Lesser flamingos (Phoenicopterus minor). Lake Bogoria, Kenya. Picture taken during filming for BBC "Life" TV Series, May 2008
Anup Shah / Impala herd (Aepyceros melampus) huddled together in the rain, Masai Mara National Reserve, Kenya. April
Daniel Heuclin / Andean flamingos / Pariwana (Phoenicopterus andinus) taking off / flying over Lake Colorada, with reflections. National Andean Fauna Reserve, Eduardo Abaroa, Bolivia, South America
Alex Mustard / Southern Stingrays (Dasyatis americana) swimming in a school over sand ripples, Grand Cayman, Cayman Islands. British West Indies. March
Jeff Rotman / Bat rays (Myliobatis californica) schooling, Redondo Rock, Galapagos Is
Tui De Roy / Red-footed booby (Sula sula), pair looking in same direction. Genovesa Island, Galapagos.
Tui De Roy / Red-footed booby (Sula sula), pair looking in same direction whilst perched in tree. Genovesa Island, Galapagos.
Tui De Roy / Red-footed booby (Sula sula), pair looking in same direction on nest with egg. Genovesa Island, Galapagos.
Tony Heald / King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September.
Alex Mustard / RF - School of blackfin barracuda (Sphyraena qenie) in open water off the wall of Yolanda Reef, Ras Mohammed Marine Park, Egypt. Red Sea. (This image may be licensed either as rights managed or roya...
Ingo Arndt / Green tree ant (Oecophylla smaragdina) group building nest by pulling on leaves together, Kakadu National Park Northern Territory, Australia. Commended in the GDT European Wildlife Photographer of th...
Ingo Arndt / Knot in foreground (Calidris canutus) and Oystercatcher, flock at high tide roost, Snettisham, RSPB Reserve, Norfolk, England, UK. November 2013.
Ingo Arndt / Migratory Locust (Locusta migratoria capito) swarm flying, near Isalo National Park, Madagascar. August 2013.