Remi Masson / European beaver (Castor fiber) swimming underwater and urinating, River Rhone, Rhone-Alpes, France, June.

Anup Shah / Forest buffalo (Syncerus caffer nanus) standing and urinating in a river running through Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Giraffe suckling with male testing urine of female Giraffa camelopardalis} Kenya

Ashley Cooper / Crabeater seal (Lobodon carcinophaga) urinating on ice with one eye shut. Antarctica. February 2019.

SCOTLAND: The Big Picture / Red Deer (Cervus elaphus) covering itself in mud from a muddy puddle after urinating in it in order to smell strong and warn off rivals, Scotland, UK, October.

Kristel Richard / Leopard (Panthera pardus) female scratching ground after urinating, South Luangwa National Park, Zambia. August

John Cancalosi / Red deer (Cervus elaphus), stag during rut, thrashing through grass and urinating on itself, England, UK. September.

Bruno D'Amicis / Wild Apennine wolf (Canis lupus italicus) adults moving on snowy mountain slope and marking a bush with their urine. Central Apennines, Abruzzo, Italy. February.. Italian endemic subspecies.

Michel Roggo / North American Beaver (Castor canadensis), urinating whilst swimming in the pool around his lodge, Northern Rockies, British Columbia, Canada July. Photographed for The Freshwater Project

Philippe Clement / Red deer (Cervus elaphus) stag with erect penis urinating on itself, rubbing antlers in grass and flicking tongue during the rut, Jaegersborg Dyrehaven, Denmark, October.

Philippe Clement / Grey wolf (Canis lupus) urinating on leftover prey, Bavarian Forest National Park, Germany, November. Captive.

Sean Viljoen / Male African wild dog (Lycaon pictus) scent marking, Khwai River, Moremi Game Reserve, Botswana.