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Ural owl (Strix uralensis) perched in tree sleeping, viewed through a fence, Estonia.
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Sven Zacek / Ural owl (Strix uralensis) perched in tree sleeping, viewed through a fence, Estonia.
School of Munk's devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacif...
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Franco Banfi / School of Munk's devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacif...
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.
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Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.
Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
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Stephane Granzotto / Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
Alpine ibex (Capra ibex) feeding in deep snow, digging to reach food, Rhone-Alpes, France, December.
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Stephane Granzotto / Alpine ibex (Capra ibex) feeding in deep snow, digging to reach food, Rhone-Alpes, France, December.
Tracking shot of a pod of Euphrosyne dolphins (Stenella coeruleoalba) swimming, Mediterranean Sea, August.
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Stephane Granzotto / Tracking shot of a pod of Euphrosyne dolphins (Stenella coeruleoalba) swimming, Mediterranean Sea, August.
Humpback Whale (Megaptera novaeangliae) beginning a deep dive. Hawaii.
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David Fleetham / Humpback Whale (Megaptera novaeangliae) beginning a deep dive. Hawaii.
Fire Dartfish / Fire Goby (Nemateleotris magnifica) against colourful coral. Fiji's outer reef.
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David Fleetham / Fire Dartfish / Fire Goby (Nemateleotris magnifica) against colourful coral. Fiji's outer reef.
Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.
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David Fleetham / Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.
Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.
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David Fleetham / Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.
Patterns of diffused sunlight on Humpback whale (Megaptera novaeangliae), Hawaii.
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David Fleetham / Patterns of diffused sunlight on Humpback whale (Megaptera novaeangliae), Hawaii.
Curious Humpback whale calf (Megaptera novaeangliae) during moment away from its mother, Hawaii.
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David Fleetham / Curious Humpback whale calf (Megaptera novaeangliae) during moment away from its mother, Hawaii.
Schooling Lemon / Milletseed butterflyfish (Chaetodon miliaris) below kayakers off Maui, Hawaii.
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David Fleetham / Schooling Lemon / Milletseed butterflyfish (Chaetodon miliaris) below kayakers off Maui, Hawaii.
Lemon sharks (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Lemon sharks (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
Lemon shark (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Lemon shark (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
Schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
Looking upwards to schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Looking upwards to schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA.
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David Fleetham / Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA.
Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.
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David Fleetham / Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.
False killer whale (Pseudorca crassidens) just below surface exhaling as it breaks the surface. Maui, Hawaii.
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David Fleetham / False killer whale (Pseudorca crassidens) just below surface exhaling as it breaks the surface. Maui, Hawaii.
Tiny Whitespotted bamboo shark (Chiloscyllium plagiosum) formed inside egg casing, Philippines.
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David Fleetham / Tiny Whitespotted bamboo shark (Chiloscyllium plagiosum) formed inside egg casing, Philippines.
Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.
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David Fleetham / Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.
Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
Schooling Brown striped snapper (Xenocys jessiae) and Yellowtail sawtail (Prionurus laticlavius), Galapagos Islands, Ecuador.
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David Fleetham / Schooling Brown striped snapper (Xenocys jessiae) and Yellowtail sawtail (Prionurus laticlavius), Galapagos Islands, Ecuador.
White mouthed moray eel (Gymnothorax meleagris), Hawaii.
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David Fleetham / White mouthed moray eel (Gymnothorax meleagris), Hawaii.
Pair of Yellow edged moray eels (Gymnothorax flavimarginatus) in coral crevice, Hawaii.
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David Fleetham / Pair of Yellow edged moray eels (Gymnothorax flavimarginatus) in coral crevice, Hawaii.
Clown anemonefish (Amphiprion ocellaris), Philippines.
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David Fleetham / Clown anemonefish (Amphiprion ocellaris), Philippines.
White-eyed moray eel (Muraena thyrsoides) close to Zebra lionfish (Dendrochirus zebra), Philippines.
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David Fleetham / White-eyed moray eel (Muraena thyrsoides) close to Zebra lionfish (Dendrochirus zebra), Philippines.
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines.
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David Fleetham / Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines.
Pair of Tomato anemonefish (Amphiprion frenatus) in Bulb tenacle sea anemone (Entacmaea quadricolor), Philippines.
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David Fleetham / Pair of Tomato anemonefish (Amphiprion frenatus) in Bulb tenacle sea anemone (Entacmaea quadricolor), Philippines.
Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.
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David Fleetham / Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.
Sargasum frogfish (Histiro histiro) near the surface floating on discarded polypropylene rope, Hawaii.
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David Fleetham / Sargasum frogfish (Histiro histiro) near the surface floating on discarded polypropylene rope, Hawaii.
Pair of White-eyed moray eels (Muraena thyrsoides) share the hole of a mooring block off Bali, Indonesia.
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David Fleetham / Pair of White-eyed moray eels (Muraena thyrsoides) share the hole of a mooring block off Bali, Indonesia.
Goby (Eviota sp) and juvenile porcelain crab (Pteroeides sp), on a Seapen. Komodo National Park, Indonesia.
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David Fleetham / Goby (Eviota sp) and juvenile porcelain crab (Pteroeides sp), on a Seapen. Komodo National Park, Indonesia.
Pixy hawkfish (Cirrhitichthys oxycephalus) in area of rich coral growth. Bali, Indonesia.
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David Fleetham / Pixy hawkfish (Cirrhitichthys oxycephalus) in area of rich coral growth. Bali, Indonesia.
Komodo dragon (Varanus komodoensis), the world's largest lizard. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), the world's largest lizard. Rinca Island, Komodo National Park, Indonesia.
Komodo dragon (Varanus komodoensis), with sailing-boat in background. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), with sailing-boat in background. Rinca Island, Komodo National Park, Indonesia.
Florida Manatee (Trichechus manatus latirostris), Three Sisters Spring, Crystal River, Florida, USA. National Wildlife Federation Photo Contest Grand Prize WINNER 2010.
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David Fleetham / Florida Manatee (Trichechus manatus latirostris), Three Sisters Spring, Crystal River, Florida, USA. National Wildlife Federation Photo Contest Grand Prize WINNER 2010.
Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
Galapagos shark (Carcharhinus galapagensis) group feeding at surface, Hawaii.
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David Fleetham / Galapagos shark (Carcharhinus galapagensis) group feeding at surface, Hawaii.
Great white shark (Carcharodon carcharias) off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) off Guadalupe Island, Mexico.
Great white shark (Carcharodon carcharias), portrait. Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias), portrait. Guadalupe Island, Mexico.
Guineafowl pufferfish (Arothron meleagris) puffed up in defence, Hawaii.
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David Fleetham / Guineafowl pufferfish (Arothron meleagris) puffed up in defence, Hawaii.
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
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Alex Mustard / American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December
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Ingo Arndt / Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December
Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December
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Ingo Arndt / Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December
Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September
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Ingo Arndt / Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September
Glaucous winged gull (Larus glaucescens) calling, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Glaucous winged gull (Larus glaucescens) calling, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) two cubs resting, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) two cubs resting, Lake Clark National Park, Alaska, USA, August
King Penguin (Aptenodytes patagonicus) rookery crowded with nesting birds incubating eggs or protecting their small chicks, Salisbury Plain, South Georgia Island, February 2011
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) rookery crowded with nesting birds incubating eggs or protecting their small chicks, Salisbury Plain, South Georgia Island, February 2011
Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 1/9
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Ingo Arndt / Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 1/9
Kingfisher (Alcedo atthis) leaving water after successfully catching a fish, Hessen, Germany, sequence 9/9
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Ingo Arndt / Kingfisher (Alcedo atthis) leaving water after successfully catching a fish, Hessen, Germany, sequence 9/9
Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 7/9
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Ingo Arndt / Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 7/9
King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February
Yellow-footed rock wallaby (Petrogale xanthopus)  Australia, December
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Ingo Arndt / Yellow-footed rock wallaby (Petrogale xanthopus) Australia, December
Intermediate Egret (Egretta intermedia) Kakadu National Park, Northern Territory, Australia, December
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Ingo Arndt / Intermediate Egret (Egretta intermedia) Kakadu National Park, Northern Territory, Australia, December
Gentoo Penguin (Pygoscelis papua) sleeping on beach, Volunteer Point, Falkland Islands, March
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Ingo Arndt / Gentoo Penguin (Pygoscelis papua) sleeping on beach, Volunteer Point, Falkland Islands, March
Antarctic Fur Seal (Arctocephalus gazella) in old Whaling Station, South Georgia, February
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) in old Whaling Station, South Georgia, February
Gentoo Penguin (Pygoscelis papua) preening, Prion Island, South Georgia Island, February
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Ingo Arndt / Gentoo Penguin (Pygoscelis papua) preening, Prion Island, South Georgia Island, February
Antarctic Fur Seal (Arctocephalus gazella) pup blonde morph, South Georgia, February
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) pup blonde morph, South Georgia, February
King Penguin (Aptenodytes patagonicus) young chick, Jason Harbour, South Georgia Island, February
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) young chick, Jason Harbour, South Georgia Island, February
King Penguins (Aptenodytes patagonicus) dispute while coming ashore, Salisbury Plain, South Georgia Island, February
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Ingo Arndt / King Penguins (Aptenodytes patagonicus) dispute while coming ashore, Salisbury Plain, South Georgia Island, February
Reindeer (Rangifer tarandus) bull, Jason Harbour, South Georgia, February
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Ingo Arndt / Reindeer (Rangifer tarandus) bull, Jason Harbour, South Georgia, February
Christmas Island Frigatebird (Fregata andrewsi) in flight, This bird is critically endangered, and endemic to Christmas Island. Christmas Island, Indian Ocean, Australian Territory. November.
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Ingo Arndt / Christmas Island Frigatebird (Fregata andrewsi) in flight, This bird is critically endangered, and endemic to Christmas Island. Christmas Island, Indian Ocean, Australian Territory. November.
Brown Booby (Sula leucogaster plotus) head portrait, with wide bill open, Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / Brown Booby (Sula leucogaster plotus) head portrait, with wide bill open, Christmas Island, Indian Ocean, Australian Territory, November
Golden Bosun / Tropicbird (Phaethon lepturus fulvus) in flight, endemic to Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / Golden Bosun / Tropicbird (Phaethon lepturus fulvus) in flight, endemic to Christmas Island, Indian Ocean, Australian Territory, November
Great frigatebird (Fregata minor listeri) drinking freshwater on the wing, Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / Great frigatebird (Fregata minor listeri) drinking freshwater on the wing, Christmas Island, Indian Ocean, Australian Territory, November
American Bison (Bison bison) head portrait of male scenting air, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) head portrait of male scenting air, South Dakota, USA
American Bison (Bison bison) herd grazing on a hillside, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd grazing on a hillside, South Dakota, USA
American Bison (Bison bison) herd stampeding, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd stampeding, South Dakota, USA
American Bison (Bison bison) herd stampeding, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd stampeding, South Dakota, USA
Flock of Rooks (Corvus frugilegus) roosting in tree at dusk, Hessen, Germany, Europe
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Ingo Arndt / Flock of Rooks (Corvus frugilegus) roosting in tree at dusk, Hessen, Germany, Europe
Flock of Bramblings (Fringilla montifringilla) perched together in branches. It's estimated that four  million individiuals return to roost during winter in Black Forest, Germany. February.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) perched together in branches. It's estimated that four million individiuals return to roost during winter in Black Forest, Germany. February.
Flock of Bramblings (Fringilla montifringilla) roosting together in branches. It's estimated that four  million individiuals roost in the Black Forest, during winter months, Germany. February.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) roosting together in branches. It's estimated that four million individiuals roost in the Black Forest, during winter months, Germany. February.
Flock of Bramblings (Fringilla montifringilla) taking off / landing on branches. It's estimated that four  million individiuals roost in the Black Forest, during winter months, Germany. February.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) taking off / landing on branches. It's estimated that four million individiuals roost in the Black Forest, during winter months, Germany. February.
Red-footed Booby (Sula sula rubripes) in flight, returning to breeding colony after sunset, Christmas Island, Indian Ocean, Australian Territory, November.
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Ingo Arndt / Red-footed Booby (Sula sula rubripes) in flight, returning to breeding colony after sunset, Christmas Island, Indian Ocean, Australian Territory, November.
Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
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Ingo Arndt / Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
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Ingo Arndt / Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
Antarctic Fur Seal (Arctocephalus gazella) bull on land, Stromness Bay, South Georgia Island, Southern Ocean, Antarctic Convergence, November 2008
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) bull on land, Stromness Bay, South Georgia Island, Southern Ocean, Antarctic Convergence, November 2008
Flock of Brambling (Fringilla montifringilla) perched at roosting tree, overwintering, Steiermark, Austria
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Ingo Arndt / Flock of Brambling (Fringilla montifringilla) perched at roosting tree, overwintering, Steiermark, Austria
Mass of Sea trout (Salmo trutta trutta) eggs, with embryos visible, Europe.
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Ingo Arndt / Mass of Sea trout (Salmo trutta trutta) eggs, with embryos visible, Europe.
Shoal of Pacific / South african sardines (Sardinops sagax) Monterey Bay Aquarium,  USA
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Ingo Arndt / Shoal of Pacific / South african sardines (Sardinops sagax) Monterey Bay Aquarium, USA
Sea trout (Salmo trutta trutta) alevin just hatched with full yolk sac, Europe
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Ingo Arndt / Sea trout (Salmo trutta trutta) alevin just hatched with full yolk sac, Europe
King penguin (Aptenodytes patagonicus) colony with adults and chicks. St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence
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Ingo Arndt / King penguin (Aptenodytes patagonicus) colony with adults and chicks. St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence
Common toad (Bufo bufo) eggs close up.
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Ingo Arndt / Common toad (Bufo bufo) eggs close up.
Flock of Bramblings (Fringilla montifringilla) in flight, an estimated four million return to roost  overwinter in Black Forest, Germany, March.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) in flight, an estimated four million return to roost overwinter in Black Forest, Germany, March.
Cape Gannet (Sula capensis) perched on a rock whilst calling, nesting colony, Lamberts Bay, South Africa.
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Ingo Arndt / Cape Gannet (Sula capensis) perched on a rock whilst calling, nesting colony, Lamberts Bay, South Africa.
King Penguin (Aptenodytes patagonicus) plumage detail, St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence.
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) plumage detail, St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence.