Sven Zacek / Ural owl (Strix uralensis) perched in tree sleeping, viewed through a fence, Estonia.
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Franco Banfi / School of Munk's devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacif...
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
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Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.
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Stephane Granzotto / Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
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Stephane Granzotto / Alpine ibex (Capra ibex) feeding in deep snow, digging to reach food, Rhone-Alpes, France, December.
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Stephane Granzotto / Tracking shot of a pod of Euphrosyne dolphins (Stenella coeruleoalba) swimming, Mediterranean Sea, August.
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David Fleetham / Humpback Whale (Megaptera novaeangliae) beginning a deep dive. Hawaii.
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David Fleetham / Fire Dartfish / Fire Goby (Nemateleotris magnifica) against colourful coral. Fiji's outer reef.
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David Fleetham / Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.
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David Fleetham / Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.
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David Fleetham / Patterns of diffused sunlight on Humpback whale (Megaptera novaeangliae), Hawaii.
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David Fleetham / Curious Humpback whale calf (Megaptera novaeangliae) during moment away from its mother, Hawaii.
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David Fleetham / Schooling Lemon / Milletseed butterflyfish (Chaetodon miliaris) below kayakers off Maui, Hawaii.
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David Fleetham / Lemon sharks (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Lemon shark (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Looking upwards to schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA.
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David Fleetham / Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.
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David Fleetham / False killer whale (Pseudorca crassidens) just below surface exhaling as it breaks the surface. Maui, Hawaii.
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David Fleetham / Tiny Whitespotted bamboo shark (Chiloscyllium plagiosum) formed inside egg casing, Philippines.
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David Fleetham / Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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David Fleetham / Schooling Brown striped snapper (Xenocys jessiae) and Yellowtail sawtail (Prionurus laticlavius), Galapagos Islands, Ecuador.
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David Fleetham / White mouthed moray eel (Gymnothorax meleagris), Hawaii.
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David Fleetham / Pair of Yellow edged moray eels (Gymnothorax flavimarginatus) in coral crevice, Hawaii.
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David Fleetham / Clown anemonefish (Amphiprion ocellaris), Philippines.
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David Fleetham / White-eyed moray eel (Muraena thyrsoides) close to Zebra lionfish (Dendrochirus zebra), Philippines.
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David Fleetham / Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines.
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David Fleetham / Pair of Tomato anemonefish (Amphiprion frenatus) in Bulb tenacle sea anemone (Entacmaea quadricolor), Philippines.
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David Fleetham / Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.
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David Fleetham / Sargasum frogfish (Histiro histiro) near the surface floating on discarded polypropylene rope, Hawaii.
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David Fleetham / Pair of White-eyed moray eels (Muraena thyrsoides) share the hole of a mooring block off Bali, Indonesia.
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David Fleetham / Goby (Eviota sp) and juvenile porcelain crab (Pteroeides sp), on a Seapen. Komodo National Park, Indonesia.
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David Fleetham / Pixy hawkfish (Cirrhitichthys oxycephalus) in area of rich coral growth. Bali, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), the world's largest lizard. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), with sailing-boat in background. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Florida Manatee (Trichechus manatus latirostris), Three Sisters Spring, Crystal River, Florida, USA. National Wildlife Federation Photo Contest Grand Prize WINNER 2010.
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David Fleetham / Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
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David Fleetham / Galapagos shark (Carcharhinus galapagensis) group feeding at surface, Hawaii.
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David Fleetham / Great white shark (Carcharodon carcharias) off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias), portrait. Guadalupe Island, Mexico.
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David Fleetham / Guineafowl pufferfish (Arothron meleagris) puffed up in defence, Hawaii.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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Alex Mustard / American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
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Ingo Arndt / Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December
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Ingo Arndt / Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December
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Ingo Arndt / Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September
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Ingo Arndt / Glaucous winged gull (Larus glaucescens) calling, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) two cubs resting, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) rookery crowded with nesting birds incubating eggs or protecting their small chicks, Salisbury Plain, South Georgia Island, February 2011
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Ingo Arndt / Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 1/9
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Ingo Arndt / Kingfisher (Alcedo atthis) leaving water after successfully catching a fish, Hessen, Germany, sequence 9/9
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Ingo Arndt / Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 7/9
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February
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Ingo Arndt / Yellow-footed rock wallaby (Petrogale xanthopus) Australia, December
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Ingo Arndt / Intermediate Egret (Egretta intermedia) Kakadu National Park, Northern Territory, Australia, December
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Ingo Arndt / Gentoo Penguin (Pygoscelis papua) sleeping on beach, Volunteer Point, Falkland Islands, March
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) in old Whaling Station, South Georgia, February
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Ingo Arndt / Gentoo Penguin (Pygoscelis papua) preening, Prion Island, South Georgia Island, February
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) pup blonde morph, South Georgia, February
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) young chick, Jason Harbour, South Georgia Island, February
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Ingo Arndt / King Penguins (Aptenodytes patagonicus) dispute while coming ashore, Salisbury Plain, South Georgia Island, February
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Ingo Arndt / Reindeer (Rangifer tarandus) bull, Jason Harbour, South Georgia, February
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Ingo Arndt / Christmas Island Frigatebird (Fregata andrewsi) in flight, This bird is critically endangered, and endemic to Christmas Island. Christmas Island, Indian Ocean, Australian Territory. November.
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Ingo Arndt / Brown Booby (Sula leucogaster plotus) head portrait, with wide bill open, Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / Golden Bosun / Tropicbird (Phaethon lepturus fulvus) in flight, endemic to Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / Great frigatebird (Fregata minor listeri) drinking freshwater on the wing, Christmas Island, Indian Ocean, Australian Territory, November
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Ingo Arndt / American Bison (Bison bison) head portrait of male scenting air, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd grazing on a hillside, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd stampeding, South Dakota, USA
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Ingo Arndt / American Bison (Bison bison) herd stampeding, South Dakota, USA
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Ingo Arndt / Flock of Rooks (Corvus frugilegus) roosting in tree at dusk, Hessen, Germany, Europe
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) perched together in branches. It's estimated that four million individiuals return to roost during winter in Black Forest, Germany. February.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) roosting together in branches. It's estimated that four million individiuals roost in the Black Forest, during winter months, Germany. February.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) taking off / landing on branches. It's estimated that four million individiuals roost in the Black Forest, during winter months, Germany. February.
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Ingo Arndt / Red-footed Booby (Sula sula rubripes) in flight, returning to breeding colony after sunset, Christmas Island, Indian Ocean, Australian Territory, November.
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Ingo Arndt / Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
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Ingo Arndt / Fallow Deer (Dama dama) at sunrise, in mist covered landscape, Moenchbruch, Germany
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Ingo Arndt / Antarctic Fur Seal (Arctocephalus gazella) bull on land, Stromness Bay, South Georgia Island, Southern Ocean, Antarctic Convergence, November 2008
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Ingo Arndt / Flock of Brambling (Fringilla montifringilla) perched at roosting tree, overwintering, Steiermark, Austria
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Ingo Arndt / Mass of Sea trout (Salmo trutta trutta) eggs, with embryos visible, Europe.
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Ingo Arndt / Shoal of Pacific / South african sardines (Sardinops sagax) Monterey Bay Aquarium, USA
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Ingo Arndt / Sea trout (Salmo trutta trutta) alevin just hatched with full yolk sac, Europe
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Ingo Arndt / King penguin (Aptenodytes patagonicus) colony with adults and chicks. St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence
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Ingo Arndt / Common toad (Bufo bufo) eggs close up.
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Ingo Arndt / Flock of Bramblings (Fringilla montifringilla) in flight, an estimated four million return to roost overwinter in Black Forest, Germany, March.
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Ingo Arndt / Cape Gannet (Sula capensis) perched on a rock whilst calling, nesting colony, Lamberts Bay, South Africa.
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Ingo Arndt / King Penguin (Aptenodytes patagonicus) plumage detail, St Andrews Bay, South Georgia Island, Southern Ocean, Antarctic Convergence.