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    • Vertebrates
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Asiatic lion (Panthera leo persica), female and cubs in morning light, cub biting mother's leg. Gir National Park, Gujarat, India. Photo Phillip Ross/Felis Images
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Felis Images / Asiatic lion (Panthera leo persica), female and cubs in morning light, cub biting mother's leg. Gir National Park, Gujarat, India. Photo Phillip Ross/Felis Images
Grey seal (Halichoerus grypus) female in surf, Horsey, Norfolk, England, UK, February.
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David Tipling / Grey seal (Halichoerus grypus) female in surf, Horsey, Norfolk, England, UK, February.
Grey seal (Halichoerus grypus) pup, Horsey, Norfolk, England, UK, February.
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David Tipling / Grey seal (Halichoerus grypus) pup, Horsey, Norfolk, England, UK, February.
Humpback whale (Megaptera novaeangliae australis) male calf playing at ocean surface after coming up to breathe. Vava'u Islands, Tonga.
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Tony Wu / Humpback whale (Megaptera novaeangliae australis) male calf playing at ocean surface after coming up to breathe. Vava'u Islands, Tonga.
Burrowing owl (Athene cunicularia) female and ten chicks aged a few weeks. Marana, Arizona, USA. May.
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Jack Dykinga / Burrowing owl (Athene cunicularia) female and ten chicks aged a few weeks. Marana, Arizona, USA. May.
Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.
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Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.
Caribbean reef shark (Carcharhinus perezi), two from below, one close up. Caribbean Sea off Gardens of the Queen National Park, Cuba.
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Shane Gross / Caribbean reef shark (Carcharhinus perezi), two from below, one close up. Caribbean Sea off Gardens of the Queen National Park, Cuba.
Ural owl (Strix uralensis) perched in tree sleeping, viewed through a fence, Estonia.
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Sven Zacek / Ural owl (Strix uralensis) perched in tree sleeping, viewed through a fence, Estonia.
Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. ??December 2016.
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Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. ??December 2016.
Glaucous Gull  (Larus hyperboreus) taking off from ice, Svalbard  Norway, July.
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Tony Heald / Glaucous Gull (Larus hyperboreus) taking off from ice, Svalbard Norway, July.
Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.
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George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.
Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.
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George Sanker / Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.
Elephant (Loxodonta africana) babies playing, Etosha National Park,Namibia.
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Tony Heald / Elephant (Loxodonta africana) babies playing, Etosha National Park,Namibia.
Ringed-tailed lemur (Lemur catta) amongst granite rocks in cave. Lemurs lick minerals from the rocks. Anja Community Reserve, Madagascar.
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Cyril Ruoso / Ringed-tailed lemur (Lemur catta) amongst granite rocks in cave. Lemurs lick minerals from the rocks. Anja Community Reserve, Madagascar.
African bush elephant, (Loxodonta africana) two head to head playing, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
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Staffan Widstrand / African bush elephant, (Loxodonta africana) two head to head playing, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Southern Giraffe (Giraffa camelopardalis) two, with one in foreground, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
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Staffan Widstrand / Southern Giraffe (Giraffa camelopardalis) two, with one in foreground, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Tawny eagle (Aquila rapax) in flight, about to land, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
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Staffan Widstrand / Tawny eagle (Aquila rapax) in flight, about to land, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Ural owl (Strix uralensis) hunting, Tartumaa county, Southern Estonia.
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Sven Zacek / Ural owl (Strix uralensis) hunting, Tartumaa county, Southern Estonia.
Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
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Oscar Dewhurst / Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
Ural owl (Strix uralensis) in flight, hunting, Tartumaa county, Southern Estonia. February.
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Sven Zacek / Ural owl (Strix uralensis) in flight, hunting, Tartumaa county, Southern Estonia. February.
Coquerel's sifaka (Propithecus coquereli) in tree, relaxing, Ankarafantsika National Park, Madagascar
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Bernard Castelein / Coquerel's sifaka (Propithecus coquereli) in tree, relaxing, Ankarafantsika National Park, Madagascar
Silky sifaka  (Propithecus candidus) in tree, Marojejy National Park, Madagascar. Critically endagered species.
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Bernard Castelein / Silky sifaka (Propithecus candidus) in tree, Marojejy National Park, Madagascar. Critically endagered species.
Giant panda (Ailuropoda melanoleuca) portrait, in falling snow. Captive bred, in enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008.
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Heather Angel / Giant panda (Ailuropoda melanoleuca) portrait, in falling snow. Captive bred, in enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008.
Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December
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Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December
Red deer (Cervus elaphus) stag roaring, silhouetted  under trees at sunrise. Bushy Park, London, UK. September
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Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September
Bengal Tiger (Panthera tigris) family drinking from pool, Ranthambhore, India
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Andy Rouse / Bengal Tiger (Panthera tigris) family drinking from pool, Ranthambhore, India
Dugong or Sea cow (Dugong dugon) split level view, Calauit Island, off Busuanga, Calamian Islands, Palawan, Philippines.
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Doug Perrine / Dugong or Sea cow (Dugong dugon) split level view, Calauit Island, off Busuanga, Calamian Islands, Palawan, Philippines.
Bighorm sheep  (Ovis canadensis) males grouped together before the rutting season. Valley of Fire State Park, Nevada, USA, February.
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Jack Dykinga / Bighorm sheep (Ovis canadensis) males grouped together before the rutting season. Valley of Fire State Park, Nevada, USA, February.
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica.
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Jordi Chias / Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica.
Humpback whale (Megaptera novaeangliae), Antarctic Peninsula, Antarctica.
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Jordi Chias / Humpback whale (Megaptera novaeangliae), Antarctic Peninsula, Antarctica.
Antarctic fur seal (Arctocephalus gazella), Antarctic Peninsula, Antarctica.
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Jordi Chias / Antarctic fur seal (Arctocephalus gazella), Antarctic Peninsula, Antarctica.
Leopard seal (Hydrurga leptonyx) resting over an iceberg, Antarctic Peninsula, Antarctica.
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Jordi Chias / Leopard seal (Hydrurga leptonyx) resting over an iceberg, Antarctic Peninsula, Antarctica.
Crabeater seal (Lobodon carcinophaga) hunting under water, Antarctic Peninsula, Antarctica.
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Jordi Chias / Crabeater seal (Lobodon carcinophaga) hunting under water, Antarctic Peninsula, Antarctica.
Antarctic fur seal (Arctocephalus gazella), Antarctic Peninsula, Antarctica.
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Jordi Chias / Antarctic fur seal (Arctocephalus gazella), Antarctic Peninsula, Antarctica.
Gentoo penguins (Pygoscelis papua) swimming together and porpoising in search of krill, Antarctic Peninsula, Antarctica.
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Jordi Chias / Gentoo penguins (Pygoscelis papua) swimming together and porpoising in search of krill, Antarctic Peninsula, Antarctica.
Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.
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Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.
Weddell seal (Leptonychotes weddellii) hauled out on ice,  Antarctic Peninsula, Antarctica.
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Jordi Chias / Weddell seal (Leptonychotes weddellii) hauled out on ice, Antarctic Peninsula, Antarctica.
Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.
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Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.
Humpback whale (Megaptera novaeangliae) spyhoping, Antarctic Peninsula, Antarctica.
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Jordi Chias / Humpback whale (Megaptera novaeangliae) spyhoping, Antarctic Peninsula, Antarctica.
Humpback whale (Megaptera novaeangliae) surfacing with baby and blowing, Antarctic Peninsula, Antarctica.
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Jordi Chias / Humpback whale (Megaptera novaeangliae) surfacing with baby and blowing, Antarctic Peninsula, Antarctica.
School of Munk's devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacif...
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Franco Banfi / School of Munk's devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacif...
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.
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Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.
Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
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Stephane Granzotto / Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
Alpine ibex (Capra ibex) feeding in deep snow, digging to reach food, Rhone-Alpes, France, December.
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Stephane Granzotto / Alpine ibex (Capra ibex) feeding in deep snow, digging to reach food, Rhone-Alpes, France, December.
Tracking shot of a pod of Euphrosyne dolphins (Stenella coeruleoalba) swimming, Mediterranean Sea, August.
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Stephane Granzotto / Tracking shot of a pod of Euphrosyne dolphins (Stenella coeruleoalba) swimming, Mediterranean Sea, August.
Humpback Whale (Megaptera novaeangliae) beginning a deep dive. Hawaii.
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David Fleetham / Humpback Whale (Megaptera novaeangliae) beginning a deep dive. Hawaii.
Fire Dartfish / Fire Goby (Nemateleotris magnifica) against colourful coral. Fiji's outer reef.
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David Fleetham / Fire Dartfish / Fire Goby (Nemateleotris magnifica) against colourful coral. Fiji's outer reef.
Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.
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David Fleetham / Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.
Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.
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David Fleetham / Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.
Patterns of diffused sunlight on Humpback whale (Megaptera novaeangliae), Hawaii.
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David Fleetham / Patterns of diffused sunlight on Humpback whale (Megaptera novaeangliae), Hawaii.
Curious Humpback whale calf (Megaptera novaeangliae) during moment away from its mother, Hawaii.
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David Fleetham / Curious Humpback whale calf (Megaptera novaeangliae) during moment away from its mother, Hawaii.
Schooling Lemon / Milletseed butterflyfish (Chaetodon miliaris) below kayakers off Maui, Hawaii.
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David Fleetham / Schooling Lemon / Milletseed butterflyfish (Chaetodon miliaris) below kayakers off Maui, Hawaii.
Lemon sharks (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Lemon sharks (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
Lemon shark (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
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David Fleetham / Lemon shark (Negaprion brevirostris) viewed from below. West End, Grand Bahamas, Caribbean.
Schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
Looking upwards to schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
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David Fleetham / Looking upwards to schooling Lemon butterflyfish (Chaetodon miliaris) near to surface. Maui, Hawaii.
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA.
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David Fleetham / Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA.
Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.
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David Fleetham / Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.
False killer whale (Pseudorca crassidens) just below surface exhaling as it breaks the surface. Maui, Hawaii.
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David Fleetham / False killer whale (Pseudorca crassidens) just below surface exhaling as it breaks the surface. Maui, Hawaii.
Tiny Whitespotted bamboo shark (Chiloscyllium plagiosum) formed inside egg casing, Philippines.
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David Fleetham / Tiny Whitespotted bamboo shark (Chiloscyllium plagiosum) formed inside egg casing, Philippines.
Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.
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David Fleetham / Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.
Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
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David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
Schooling Brown striped snapper (Xenocys jessiae) and Yellowtail sawtail (Prionurus laticlavius), Galapagos Islands, Ecuador.
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David Fleetham / Schooling Brown striped snapper (Xenocys jessiae) and Yellowtail sawtail (Prionurus laticlavius), Galapagos Islands, Ecuador.
White mouthed moray eel (Gymnothorax meleagris), Hawaii.
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David Fleetham / White mouthed moray eel (Gymnothorax meleagris), Hawaii.
Pair of Yellow edged moray eels (Gymnothorax flavimarginatus) in coral crevice, Hawaii.
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David Fleetham / Pair of Yellow edged moray eels (Gymnothorax flavimarginatus) in coral crevice, Hawaii.
Clown anemonefish (Amphiprion ocellaris), Philippines.
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David Fleetham / Clown anemonefish (Amphiprion ocellaris), Philippines.
White-eyed moray eel (Muraena thyrsoides) close to Zebra lionfish (Dendrochirus zebra), Philippines.
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David Fleetham / White-eyed moray eel (Muraena thyrsoides) close to Zebra lionfish (Dendrochirus zebra), Philippines.
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines.
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David Fleetham / Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines.
Pair of Tomato anemonefish (Amphiprion frenatus) in Bulb tenacle sea anemone (Entacmaea quadricolor), Philippines.
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David Fleetham / Pair of Tomato anemonefish (Amphiprion frenatus) in Bulb tenacle sea anemone (Entacmaea quadricolor), Philippines.
Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.
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David Fleetham / Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.
Sargasum frogfish (Histiro histiro) near the surface floating on discarded polypropylene rope, Hawaii.
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David Fleetham / Sargasum frogfish (Histiro histiro) near the surface floating on discarded polypropylene rope, Hawaii.
Pair of White-eyed moray eels (Muraena thyrsoides) share the hole of a mooring block off Bali, Indonesia.
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David Fleetham / Pair of White-eyed moray eels (Muraena thyrsoides) share the hole of a mooring block off Bali, Indonesia.
Goby (Eviota sp) and juvenile porcelain crab (Pteroeides sp), on a Seapen. Komodo National Park, Indonesia.
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David Fleetham / Goby (Eviota sp) and juvenile porcelain crab (Pteroeides sp), on a Seapen. Komodo National Park, Indonesia.
Pixy hawkfish (Cirrhitichthys oxycephalus) in area of rich coral growth. Bali, Indonesia.
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David Fleetham / Pixy hawkfish (Cirrhitichthys oxycephalus) in area of rich coral growth. Bali, Indonesia.
Komodo dragon (Varanus komodoensis), the world's largest lizard. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), the world's largest lizard. Rinca Island, Komodo National Park, Indonesia.
Komodo dragon (Varanus komodoensis), with sailing-boat in background. Rinca Island, Komodo National Park, Indonesia.
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David Fleetham / Komodo dragon (Varanus komodoensis), with sailing-boat in background. Rinca Island, Komodo National Park, Indonesia.
Florida Manatee (Trichechus manatus latirostris), Three Sisters Spring, Crystal River, Florida, USA. National Wildlife Federation Photo Contest Grand Prize WINNER 2010.
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David Fleetham / Florida Manatee (Trichechus manatus latirostris), Three Sisters Spring, Crystal River, Florida, USA. National Wildlife Federation Photo Contest Grand Prize WINNER 2010.
Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) with mouth open, just below the surface off Guadalupe Island, Mexico.
Galapagos shark (Carcharhinus galapagensis) group feeding at surface, Hawaii.
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David Fleetham / Galapagos shark (Carcharhinus galapagensis) group feeding at surface, Hawaii.
Great white shark (Carcharodon carcharias) off Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias) off Guadalupe Island, Mexico.
Great white shark (Carcharodon carcharias), portrait. Guadalupe Island, Mexico.
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David Fleetham / Great white shark (Carcharodon carcharias), portrait. Guadalupe Island, Mexico.
Guineafowl pufferfish (Arothron meleagris) puffed up in defence, Hawaii.
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David Fleetham / Guineafowl pufferfish (Arothron meleagris) puffed up in defence, Hawaii.
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
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David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.
American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
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Alex Mustard / American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December
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Ingo Arndt / Cape gannets (Morus capensis / Sula capensis) at nesting colony, zooming from tight to wide shot, Lamberts Bay, South Africa, December
Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December
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Ingo Arndt / Cape fur seals (Arctocephalus pusillus) at colony, Cape Cross, Namibia, December
Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September
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Ingo Arndt / Grizzly bear (Ursus arctos horribilis) mother with two cubs catching salmon, Lake Clark National Park, Alaska, USA, September
Glaucous winged gull (Larus glaucescens) calling, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Glaucous winged gull (Larus glaucescens) calling, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August
Grizzly Bear (Ursus arctos horribilis) two cubs resting, Lake Clark National Park, Alaska, USA, August
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Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) two cubs resting, Lake Clark National Park, Alaska, USA, August