Wim van den Heever / Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April.

Cheryl-Samantha Owen / Great white shark (Carcharodon carcharias) breaching, at decoy seal, Seal Island, False Bay, off the coast of Cape Town, South Africa. July.

Edwin Giesbers / Barbary macaque (Macaca sylvanus) baring teeth as a sign of submission, Gibraltar Nature Reserve, Gibraltar,

Wim van den Heever / Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Sequence 4 of 5

Sven Zacek / Otter (lutra lutra) eating Gudgeon (Gobio gobio) in Southern Estonia, January.

Staffan Widstrand / Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa. Specially commended in the Animal Portraits category of...

Staffan Widstrand / Eurasian griffon vulture (Gyps fulvus) blurred motion photograph at wildlife watching and vulture feeding site, Pre-Pyrenees, Catalonia, Spain, March.

Fabrice Cahez / Wild cat (Felis sylvestris) in snow, Vosges, France, February.

Tony Heald / Bengal tiger (Panthera tigris tigris) walking, Bandhavgarh National Park, India.

Yukihiro Fukuda / Japanese macaque (Macaca fuscata fuscata) rare white furred baby jumping, Jigokudani Valley, Nagano Prefecture, Japan. June.

Andy Rouse / Bengal tiger (Panthera tigris tigris) portrait, whilst stalking, Bandhavgarh, India.

Espen Bergersen / Common raven (Corvus corax) pair in courtship display, flying upsidedown and calling, Hornoya bird cliff, Norway. March.