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    • Viperidae
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Sumatran pit viper (Trimeresurus sumatranus) juvenile on tree trunk, in tropical rainforest. Danum Valley, Sabah, Borneo, Malaysia.
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Nick Garbutt / Sumatran pit viper (Trimeresurus sumatranus) juvenile on tree trunk, in tropical rainforest. Danum Valley, Sabah, Borneo, Malaysia.
Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April
01622435
David Tipling / Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April
01622434
David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
01622430
David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
Meadow viper, (Vipera ursinii), basking in alpine meadow, Apennines, Italy, September . Non-ex.
01632888
Fabio Pupin / Meadow viper, (Vipera ursinii), basking in alpine meadow, Apennines, Italy, September . Non-ex.
Peringuey's desert adder, (Bitis peringueyi), sidewinding on dune, Namib desert, Namibia, February . Non-ex.
01632782
Fabio Pupin / Peringuey's desert adder, (Bitis peringueyi), sidewinding on dune, Namib desert, Namibia, February . Non-ex.
Horned viper (Vipera ammodytes) in natural setting, Turkey.
01620962
Daniel Heuclin / Horned viper (Vipera ammodytes) in natural setting, Turkey.
Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
01625450
John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
Ecuadorian Toad-headed Pit-viper (Bothrocophias campbelli) in rainforest, Mashpi Amagusa Reserve, Pichincha, Ecuador
00583296
Chien Lee / Ecuadorian Toad-headed Pit-viper (Bothrocophias campbelli) in rainforest, Mashpi Amagusa Reserve, Pichincha, Ecuador
Puff Adder (Bitis arietans) near wetland, Marakele National Park, Limpopo, South Africa
00573479
Pete Oxford / Puff Adder (Bitis arietans) near wetland, Marakele National Park, Limpopo, South Africa
Peringuey's adder (Bitis peringueyi) buried in sand,  flicking tongue, hunting, Namib Desert, Namibia.
01560105
Christophe Courteau / Peringuey's adder (Bitis peringueyi) buried in sand, flicking tongue, hunting, Namib Desert, Namibia.
Western rattlesnake (Crotalus viridis viridis), threat display, South Dakota, USA, September.
01512025
Silke Arndt / Western rattlesnake (Crotalus viridis viridis), threat display, South Dakota, USA, September.
Western diamondback rattlesnake (Crotalus atrox), defensive rattling, Arizona, USA, April.
01508236
John Cancalosi / Western diamondback rattlesnake (Crotalus atrox), defensive rattling, Arizona, USA, April.
Group of gravid female Timber rattlesnakes (Crotalus horridus) basking before giving birth, Pennsylvania, USA, July.
01508163
John Cancalosi / Group of gravid female Timber rattlesnakes (Crotalus horridus) basking before giving birth, Pennsylvania, USA, July.
Female Timber rattlesnake (Crotalus horridus) at nest site with newborn young, Pennsylvania, USA, September.
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John Cancalosi / Female Timber rattlesnake (Crotalus horridus) at nest site with newborn young, Pennsylvania, USA, September.
Puff adder (Bitis arietans) moving into ambush position under a bush, DeHoop Nature Reserve, Western Cape, South Africa, January.
01475191
Tony Phelps / Puff adder (Bitis arietans) moving into ambush position under a bush, DeHoop Nature Reserve, Western Cape, South Africa, January.
Sunda island pitviper (Trimeresurus insularis) in tree, Komodo Island.
01555328
Daniel Heuclin / Sunda island pitviper (Trimeresurus insularis) in tree, Komodo Island.
Western diamond-backed rattlesnake (Crotalus atrox) Texas, USA, April.
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Bence Mate / Western diamond-backed rattlesnake (Crotalus atrox) Texas, USA, April.
Prairie rattlesnake (Crotalus viridis viridis), threat display, South Dakota, USA September. Sequence 1 of 3
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Ingo Arndt / Prairie rattlesnake (Crotalus viridis viridis), threat display, South Dakota, USA September. Sequence 1 of 3
Prairie Rattlesnake (Crotalus viridis viridis) in defensive posture, South Dakota
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Ingo Arndt / Prairie Rattlesnake (Crotalus viridis viridis) in defensive posture, South Dakota
Osborne's Lancehead (Bothrops osbornei) juvenile showing differently colored tail tip used for caudal luring, native to South America
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Pete Oxford / Osborne's Lancehead (Bothrops osbornei) juvenile showing differently colored tail tip used for caudal luring, native to South America
Fer-de-lance (Bothrops asper), native to South America
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Pete Oxford / Fer-de-lance (Bothrops asper), native to South America
Sand viper (Cerastes vipera) sidewinding on sand dunes. Grand Erg Oriental, Kebili Governorate. Tunisia.
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Bruno D'Amicis / Sand viper (Cerastes vipera) sidewinding on sand dunes. Grand Erg Oriental, Kebili Governorate. Tunisia.
Adder (Vipera berus) basking amongst bracken fronds. Peak District National Park, Derbyshire, UK. April.
01509998
Alex Hyde / Adder (Vipera berus) basking amongst bracken fronds. Peak District National Park, Derbyshire, UK. April.
Western diamondback rattlesnake (Crotalus atrox) tasting air, in habitat, Arizona, USA, October. Controlled conditions
01508335
Daniel Heuclin / Western diamondback rattlesnake (Crotalus atrox) tasting air, in habitat, Arizona, USA, October. Controlled conditions
Banded sea kraits (Laticauda colubrina) around the roots of mangrove tree, split level. Mali Island, Macuata Province, Fiji, South Pacific.
01508273
Juergen Freund / Banded sea kraits (Laticauda colubrina) around the roots of mangrove tree, split level. Mali Island, Macuata Province, Fiji, South Pacific.
Sidewinder (Crotalus cerstastes) in sand dunes, Anza-Borrego Desert State Park, California, USA, May.
01490546
Chris Mattison / Sidewinder (Crotalus cerstastes) in sand dunes, Anza-Borrego Desert State Park, California, USA, May.
Mojave rattlesnake (Crotalus scutulatus) Mojave desert, California, June.
01448577
Daniel Heuclin / Mojave rattlesnake (Crotalus scutulatus) Mojave desert, California, June.
Rhinoceros viper (Bitis nasicornis), from Central Africa, captive, July.
01448560
Daniel Heuclin / Rhinoceros viper (Bitis nasicornis), from Central Africa, captive, July.
Western diamondback rattlesnake (Crotalus atrox) Rio Grande Valley, Texas, USA, October. meetyourneighbours.net project
01376183
MYN / Seth Patterson / Western diamondback rattlesnake (Crotalus atrox) Rio Grande Valley, Texas, USA, October. meetyourneighbours.net project
Large scaled pit viper (Trimeresurus macrolepis) head profile, water droplet below, Western Ghats, Southern India
01388669
Sandesh Kadur / Large scaled pit viper (Trimeresurus macrolepis) head profile, water droplet below, Western Ghats, Southern India
Baskian viper (Vipera seoanei) Geres National Park, Portugal, September.
01365832
Luis Quinta / Baskian viper (Vipera seoanei) Geres National Park, Portugal, September.
RF- Western Diamondback Rattlesnake (Crotalus atrox), adult in striking pose with rattle raised, in wildflowers. Laredo, Webb County, South Texas, USA. April. (This image may be licensed either as rig...
01350505
Rolf Nussbaumer / RF- Western Diamondback Rattlesnake (Crotalus atrox), adult in striking pose with rattle raised, in wildflowers. Laredo, Webb County, South Texas, USA. April. (This image may be licensed either as rig...
Eastern Diamondback Rattlesnake (Crotalus adamanteus), Little St. Simon's Island, Georgia
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Pete Oxford / Eastern Diamondback Rattlesnake (Crotalus adamanteus), Little St. Simon's Island, Georgia
Adder (Vipera berus) portrait, Staffordshire, England, UK, April
01341072
Danny Green / 2020VISION / Adder (Vipera berus) portrait, Staffordshire, England, UK, April
Adder (Vipera berus) coiled, basking on moss in the spring sunshine, Staffordshire, England, UK, April. Photographer quote: "OK so snakes are perhaps not as sexy as seals or foxes but I just can't get...
01341069
Danny Green / 2020VISION / Adder (Vipera berus) coiled, basking on moss in the spring sunshine, Staffordshire, England, UK, April. Photographer quote: "OK so snakes are perhaps not as sexy as seals or foxes but I just can't get...
Adder (Vipera berus) tasting the air with tongue,  Staffordshire, England, UK, April. 2020VISION Exhibition. 2020VISION Book Plate.
01341066
Danny Green / 2020VISION / Adder (Vipera berus) tasting the air with tongue, Staffordshire, England, UK, April. 2020VISION Exhibition. 2020VISION Book Plate.
Adder (Vipera berus) head portrait, Staffordshire, England, UK, April
01341059
Danny Green / 2020VISION / Adder (Vipera berus) head portrait, Staffordshire, England, UK, April
Adder (Vipera berus) basking in the spring, Staffordshire, England, UK, April
01341056
Danny Green / 2020VISION / Adder (Vipera berus) basking in the spring, Staffordshire, England, UK, April
Adder (Vipera berus) basking in the spring sunshine flicking tongue, England, UK, April. Did you know? The Adder is the only venomous snake in the UK.
01341054
Danny Green / 2020VISION / Adder (Vipera berus) basking in the spring sunshine flicking tongue, England, UK, April. Did you know? The Adder is the only venomous snake in the UK.
Eyelash Viper (Bothriechis schlegelii) with yellow coloration, coiled on branch, Cahuita National Park, Costa Rica
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Thomas Marent / Eyelash Viper (Bothriechis schlegelii) with yellow coloration, coiled on branch, Cahuita National Park, Costa Rica
Eyelash viper {Bothrops schlegeli} on Heliconia,  Esmeraldas, Ecuador
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Pete Oxford / Eyelash viper {Bothrops schlegeli} on Heliconia, Esmeraldas, Ecuador
Eyelash viper coiled {Bothrops schlegeli} captive
01123348
LYNN M. STONE / Eyelash viper coiled {Bothrops schlegeli} captive
Eyelash Viper (Bothriechis schlegelii) venomous, arboreal, Esmeraldas, Ecuador
00217187
Pete Oxford / Eyelash Viper (Bothriechis schlegelii) venomous, arboreal, Esmeraldas, Ecuador
Colorado Desert Sidewinder (Crotalus cerastes laterorepens) on sand, venomous species native to deserts of California and Mexico
00117441
ZSSD / Colorado Desert Sidewinder (Crotalus cerastes laterorepens) on sand, venomous species native to deserts of California and Mexico
Peringuey's Sidewinding Adder (Bitis peringueyi) sidewinding across sand dune, Namib Desert, Namibia
00510523
Michael & Patricia Fogden / Peringuey's Sidewinding Adder (Bitis peringueyi) sidewinding across sand dune, Namib Desert, Namibia
Peringuey's Sidewinding Adder (Bitis peringueyi) on sand dune, Namib Desert, Namibia
00510522
Michael & Patricia Fogden / Peringuey's Sidewinding Adder (Bitis peringueyi) on sand dune, Namib Desert, Namibia
Two-striped Forest Pit Viper (Bothrops bilineatus) coiled around plant, front view, Atlantic Forest ecosystem, Brazil
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Claus Meyer / Two-striped Forest Pit Viper (Bothrops bilineatus) coiled around plant, front view, Atlantic Forest ecosystem, Brazil
Russell's Viper (Daboia russelii) being milked for venom, Madras India
00510629
Michael & Patricia Fogden / Russell's Viper (Daboia russelii) being milked for venom, Madras India
Eyelash Viper (Bothriechis schlegelii) gold morph camouflaged among palm fruits, rainforest, Costa Rica
00510923
Michael & Patricia Fogden / Eyelash Viper (Bothriechis schlegelii) gold morph camouflaged among palm fruits, rainforest, Costa Rica
Eyelash Viper (Bothriechis schlegelii) showing loreal pits for detecting heat, rainforest, Costa Rica
00510921
Michael & Patricia Fogden / Eyelash Viper (Bothriechis schlegelii) showing loreal pits for detecting heat, rainforest, Costa Rica
Eyelash Viper (Bothriechis schlegelii) gold morph lying in wait on Heliconia (Heliconia imbricata) to strike at Rufous-tailed Hummingbird (Amazilia tzacatl) rainforest, Costa Rica
00510928
Michael & Patricia Fogden / Eyelash Viper (Bothriechis schlegelii) gold morph lying in wait on Heliconia (Heliconia imbricata) to strike at Rufous-tailed Hummingbird (Amazilia tzacatl) rainforest, Costa Rica
Bushmaster (Lachesis muta) baby hatching from egg, lowland rainforest on Atlantic slope of Costa Rica
00511319
Michael & Patricia Fogden / Bushmaster (Lachesis muta) baby hatching from egg, lowland rainforest on Atlantic slope of Costa Rica
Black-tailed Rattlesnake (Crotalus molossus) portrait, Chiricahua Mountains, Arizona
00510419
Michael & Patricia Fogden / Black-tailed Rattlesnake (Crotalus molossus) portrait, Chiricahua Mountains, Arizona
Two-striped Forest Pit Viper (Bothrops bilineatus) hanging on a Heliconia (Heliconia sp) flower, Atlantic Forest, Brazil
00300224
Claus Meyer / Two-striped Forest Pit Viper (Bothrops bilineatus) hanging on a Heliconia (Heliconia sp) flower, Atlantic Forest, Brazil
Western Rattlesnake (Crotalus viridis) in defensive posture, South Dakota
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Jim Brandenburg / Western Rattlesnake (Crotalus viridis) in defensive posture, South Dakota
Red Rattlesnake (Crotalus ruber) coiled on ground, Baja California, Mexico
00105463
Larry Minden / Red Rattlesnake (Crotalus ruber) coiled on ground, Baja California, Mexico
Red Rattlesnake (Crotalus ruber) with tongue extended, Baja California, Mexico
00105462
Larry Minden / Red Rattlesnake (Crotalus ruber) with tongue extended, Baja California, Mexico
Western Rattlesnake (Crotalus viridis) in defensive posture, South Dakota
00113409
Jim Brandenburg / Western Rattlesnake (Crotalus viridis) in defensive posture, South Dakota
Eyelash viper {Bothrops schlegeli} on Heliconia Esmeraldas, Ecuador
01090702
Pete Oxford / Eyelash viper {Bothrops schlegeli} on Heliconia Esmeraldas, Ecuador
Slow motion clip of a Fer de Lance (Bothrops atrox) smelling with its tongue, Orellana Province, Ecuador.
01631726
Morley Read / Slow motion clip of a Fer de Lance (Bothrops atrox) smelling with its tongue, Orellana Province, Ecuador.
Fer de Lance (Bothrops atrox) on the rainforest floor, Orellana Province, Ecuador.
01631725
Morley Read / Fer de Lance (Bothrops atrox) on the rainforest floor, Orellana Province, Ecuador.
Timber rattlesnake (Crotalus horridus) females coiled up in leaf litter, newborn young sheltering under rock. At maternity site, Pennsylvania, USA. August.
01633587
John Cancalosi / Timber rattlesnake (Crotalus horridus) females coiled up in leaf litter, newborn young sheltering under rock. At maternity site, Pennsylvania, USA. August.
Timber rattlesnake (Crotalus horridus) females in undergrowth at maternity site, Pennsylvania, USA. August.
01633586
John Cancalosi / Timber rattlesnake (Crotalus horridus) females in undergrowth at maternity site, Pennsylvania, USA. August.
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August.
01633584
John Cancalosi / Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August.
Puff adder (Bitis arietans), Zimanga game reserve, KwaZulu-Natal, South Africa.
01633092
Ann & Steve Toon / Puff adder (Bitis arietans), Zimanga game reserve, KwaZulu-Natal, South Africa.
Milos Viper (Macrovipera schweizeri) inhabiting the rocky shoreline of a freshwater lake on Milos Island, Greece, July. Controlled conditions.
01633201
Robert Valentic / Milos Viper (Macrovipera schweizeri) inhabiting the rocky shoreline of a freshwater lake on Milos Island, Greece, July. Controlled conditions.
Milos Viper (Macrovipera schweizeri) sub adult luring potential prey to within striking range with tail. Milos Island, Greece, July. Controlled conditions.
01633200
Robert Valentic / Milos Viper (Macrovipera schweizeri) sub adult luring potential prey to within striking range with tail. Milos Island, Greece, July. Controlled conditions.
Milos Viper (Macrovipera schweizeri) sub-adult in the grounds of a church, Milos Island, Greece, July. Controlled conditions.
01633199
Robert Valentic / Milos Viper (Macrovipera schweizeri) sub-adult in the grounds of a church, Milos Island, Greece, July. Controlled conditions.
Milos Viper (Macrovipera schweizeri) from the western side of Milos Island, Greece, July. Controlled conditions.
01633198
Robert Valentic / Milos Viper (Macrovipera schweizeri) from the western side of Milos Island, Greece, July. Controlled conditions.
Milos Viper (Macrovipera schweizeri) sub-adult basking along a rock outcrop on a lake shoreline on Milos Island, Greece, July. Controlled conditions.
01633197
Robert Valentic / Milos Viper (Macrovipera schweizeri) sub-adult basking along a rock outcrop on a lake shoreline on Milos Island, Greece, July. Controlled conditions.
Panamint rattlesnake (Crotalus stephensi) with tongue out and rattle raised, on rock. Amargosa Desert, Nye County, Nevada. May.
01633204
Daniel Heuclin / Panamint rattlesnake (Crotalus stephensi) with tongue out and rattle raised, on rock. Amargosa Desert, Nye County, Nevada. May.
Panamint rattlesnake (Crotalus stephensi) on rock, Amargosa Desert, Nye County, Nevada. May.
01633203
Daniel Heuclin / Panamint rattlesnake (Crotalus stephensi) on rock, Amargosa Desert, Nye County, Nevada. May.
Panamint rattlesnake (Crotalus stephensi) on rock, tongue out and rattle raised. Amargosa Desert, Nye County, Nevada. May.
01633202
Daniel Heuclin / Panamint rattlesnake (Crotalus stephensi) on rock, tongue out and rattle raised. Amargosa Desert, Nye County, Nevada. May.
Sumatran pit viper (Trimeresurus sumatranus) juvenile on trunk. Danum Valley, Sabah, Borneo, Malaysia. Focus stacked image.
01634541
Nick Garbutt / Sumatran pit viper (Trimeresurus sumatranus) juvenile on trunk. Danum Valley, Sabah, Borneo, Malaysia. Focus stacked image.
Western diamondback rattlesnake (Crotalus atrox) portrait, coiled up in evening light. Texas, USA. June.
01633494
Karine Aigner / Western diamondback rattlesnake (Crotalus atrox) portrait, coiled up in evening light. Texas, USA. June.
Western diamondback rattlesnake (Crotalus atrox). Texas, USA. June.
01633493
Karine Aigner / Western diamondback rattlesnake (Crotalus atrox). Texas, USA. June.
Western diamondback rattlesnake (Crotalus atrox) crossing a track, Texas, USA. June.
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Karine Aigner / Western diamondback rattlesnake (Crotalus atrox) crossing a track, Texas, USA. June.
Western diamondback rattlesnake (Crotalus atrox) tail rattle. Texas, USA. May.
01633488
Karine Aigner / Western diamondback rattlesnake (Crotalus atrox) tail rattle. Texas, USA. May.
Western diamondback rattlesnake (Crotalus atrox) coiled up at base of Prickly pear cactus (Opuntia sp). Texas, USA. April.
01633462
Karine Aigner / Western diamondback rattlesnake (Crotalus atrox) coiled up at base of Prickly pear cactus (Opuntia sp). Texas, USA. April.
Western diamondback rattlesnake (Crotalus atrox) coiled up. Texas, USA. April.
01633454
Karine Aigner / Western diamondback rattlesnake (Crotalus atrox) coiled up. Texas, USA. April.
Researcher from Indio Mountain Research Station capturing Eastern black-tailed rattlesnake (Crotalus ornatus) Chihuahuan Desert,Texas, USA. May 2019.
01630557
Daniel Heuclin / Researcher from Indio Mountain Research Station capturing Eastern black-tailed rattlesnake (Crotalus ornatus) Chihuahuan Desert,Texas, USA. May 2019.
Black-tailed rattlesnake (Crotalus molossus). Chiricahua mountains, Arizona, USA. June.
01630550
Daniel Heuclin / Black-tailed rattlesnake (Crotalus molossus). Chiricahua mountains, Arizona, USA. June.
Western diamondback rattlesnake (Crotalus atrox) flicking tongue and with rattle raised. Arizona, USA. Controlled conditions.
01630547
Daniel Heuclin / Western diamondback rattlesnake (Crotalus atrox) flicking tongue and with rattle raised. Arizona, USA. Controlled conditions.
Western diamondback rattlesnake (Crotalus atrox) camouflaged amongst rocks, with rattle raised. Sonoran Desert, Arizona, USA. Controlled conditions. June 2019.
01630546
Daniel Heuclin / Western diamondback rattlesnake (Crotalus atrox) camouflaged amongst rocks, with rattle raised. Sonoran Desert, Arizona, USA. Controlled conditions. June 2019.
Western diamondback rattlesnake (Crotalus atrox) with rattle raised, Arizona, USA. Controlled conditions.
01630545
Daniel Heuclin / Western diamondback rattlesnake (Crotalus atrox) with rattle raised, Arizona, USA. Controlled conditions.
Western diamondback rattlesnake (Crotalus atrox), flicking tongue and with rattle raised, Arizona, USA. Controlled conditions.
01630543
Daniel Heuclin / Western diamondback rattlesnake (Crotalus atrox), flicking tongue and with rattle raised, Arizona, USA. Controlled conditions.
Common European adder (Vipera berus) male warming up in sun trap on heath, Holt, North Norfolk, England, UK. April.
01622438
David Tipling / Common European adder (Vipera berus) male warming up in sun trap on heath, Holt, North Norfolk, England, UK. April.
Common European adder (Vipera berus) male guarding female before mating on heath, Holt, North Norfolk, England, UK. April.
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David Tipling / Common European adder (Vipera berus) male guarding female before mating on heath, Holt, North Norfolk, England, UK. April.
Common European adder (Vipera berus) male guarding female before mating on heath Holt, North Norfolk, England, UK. April.
01622436
David Tipling / Common European adder (Vipera berus) male guarding female before mating on heath Holt, North Norfolk, England, UK. April.
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
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David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
01622432
David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
Common European adder (Vipera berus) male and female warming up in sun trap on heath, Holt, North Norfolk, England, UK. March.
01622429
David Tipling / Common European adder (Vipera berus) male and female warming up in sun trap on heath, Holt, North Norfolk, England, UK. March.
Common European adder (Vipera berus) male and female warming up in sun trap on heath, Holt, North Norfolk, England, UK. March.
01622428
David Tipling / Common European adder (Vipera berus) male and female warming up in sun trap on heath, Holt, North Norfolk, England, UK. March.
Adders (Vipera berus) emerging from hibernation to soak up the warm sunshine. Near Holt, North Norfolk, England, UK. February.
01622425
David Tipling / Adders (Vipera berus) emerging from hibernation to soak up the warm sunshine. Near Holt, North Norfolk, England, UK. February.
Walser's viper, (Vipera walser), recently described new species, adult basking in habitat, Alps, Italy, June . Non-ex.
01632919
Fabio Pupin / Walser's viper, (Vipera walser), recently described new species, adult basking in habitat, Alps, Italy, June . Non-ex.