Ismaele Tortella / White tailed eagle (Haliaeetus albicilla) catching fish, Lofoten, Norway, August.

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Hermann Brehm / Cuckoo (Cuculus canorus) in flight, Germany, April. May

Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Nick Garbutt / Black-browed albatross (Thalassarche melanophris) flying over South Atlantic. Cooper Bay, South Georgia. November.

Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.

Tony Heald / Glaucous Gull (Larus hyperboreus) taking off from ice, Svalbard Norway, July.

Staffan Widstrand / Tawny eagle (Aquila rapax) in flight, about to land, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Sven Zacek / Ural owl (Strix uralensis) hunting, Tartumaa county, Southern Estonia.

Sven Zacek / Ural owl (Strix uralensis) in flight, hunting, Tartumaa county, Southern Estonia. February.

Jordi Chias / Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica.

Terry Whittaker / Man collecting seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down cliff to collect seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being raised up cliff to collect seabird eggs, including those of Common guillemots (Uria aalge), Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Jack Dykinga / Sandhill cranes (Antigone canadensis) and Snow geese (Chen caerulescens) at sunrise, Bosque del Apache National Wildlife Refuge, New Mexico, USA, November.

Jack Dykinga / Bald eagle (Haliaeetus leucocephalus) perched as Snow geese (Chen caerulescens) fly by, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into a tight packed flock on lake, Bosque del Apache National Wildlife Refuge, New Mexico, USA. January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into flock, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Sandhill cranes (Antigone canadensis) taking off and flying past a full 'super-moon' en route to feeding in neighboring fields. Bosque del Apache National Wildlife Refuge, New Mexico, USA, Jan...

Ernie Janes / Brent geese (Branta bernicla) flying with moon, Titchwell, Norfolk, UK, December

Winfried Wisniewski / Black Kite (Milvus migrans) flying, Castile-La Mancha, Spain

John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019

John Shaw / Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May.

Andy Rouse / Barn owl (Tyto alba) hunting with sunset light on wings, UK. March.

Espen Bergersen / Killer whales / orcas (Orcinus orca). Spyhopping. Kvaloya, Troms, Norway November

Espen Bergersen / Gulls and Killer whales / orcas (Orcinus orca) feeding on herring. Kvaloya, Troms, Norway October

Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.

Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.

Andy Rouse / Kingfisher, (Alcedo atthis), diving for fish from branch, UK, Medium repro only

Andy Rouse / Kingfisher, (Alcedo atthis), diving for fish, UK, Medium repro only, Motion blur

Andy Rouse / Kingfisher, (Alcedo atthis), diving for fish, UK

Andy Rouse / Kingfisher, (Alcedo atthis), diving for fish, UK

Jack Dykinga / Sandhill crane (Antigone canadensis) in flight against blue sky, Bosque Del Apache, New Mexico, USA. December.

Jack Dykinga / Sandhill cranes (Antigone canadensis) in flight in late evening light, Bosque Del Apache, New Mexico, USA. December

Jack Dykinga / Flocks of Mallard ducks (Anas platyrhynchos) taking off in tree-lined wetlands. Bosque del Apache National Wildlife Refuge, New Mexico, USA.

Jack Dykinga / Yellow-headed blackbirds (Xanthocephalus xanthocephalus) and Redwing blackbirds (Agelaius phoeniceus) roosting. Whitewater Draw Wildlife Area, Arizona, USA, December.

Jack Dykinga / Murmuration of Yellow-headed blackbirds (Xanthocephalus xanthocephalus) and Redwing blackbirds (Agelaius phoeniceus) at sunset, before roosting. Whitewater Draw Wildlife Area, Arizona, USA, December.

SCOTLAND: The Big Picture / Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April.

Oscar Dewhurst / Great bittern (Botaurus stellaris) in flight over water. London, UK. December.

Sandesh Kadur / Slow motion clip of Amur falcons (Falco amurensis) congregating at roost site during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) hunting Atlantic herring (Clupea harengus), Troms, Norway, October.

Espen Bergersen / Aerial shot of a pod of Killer whales (Orcinus orca) carousel feeding, Skjervoy, Troms, Norway, November.

Terry Whittaker / Grey heron ( Ardea cinerea) flying to bin. Herons congregate around the fish stalls as city markets are closing, picking up scraps of food. Amsterdam, Netherlands. April.

Ross Hoddinott / Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August.

Ross Hoddinott / Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. July.

Ross Hoddinott / Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. July.

Marie Read / Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July.

Marie Read / Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August.

Marie Read / Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January.

Marie Read / Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

Konrad Wothe / Sword-billed hummingbird (Ensifera ensifera) in flight, North-Ecuador, Ecuador.

Konrad Wothe / Small tortoiseshell butterfly (Aglais urtica) in flight, with flower, Bavaria, Germany.

David Pattyn / Golden eagle (Aquila chrysaetos) in flight in front of a vertical rocky mountain side, Gran Paradiso National Park, Aosta Valley, Italy, May

Alan Murphy/ BIA / Barn Owl (Tyto alba) flying with rodent prey, Texas

Alan Murphy/ BIA / Harris' Hawk (Parabuteo unicinctus) flying, Texas

Alan Murphy/ BIA / Northern Caracara (Caracara cheriway) pair fighting, Texas

Alan Murphy/ BIA / Baltimore Oriole (Icterus galbula) males fighting, Texas

Martin Grimm/ BIA / Eurasian Kestrel (Falco tinnunculus) male flying, Baden-Wurttemberg, Germany

Andres M. Dominguez/ BIA / Common Tern (Sterna hirundo) flying with fish prey, Danube Delta, Romania

Avi Meir/ BIA / Slender-billed Gull (Larus genei) landing, Eilat, Israel

Alan Murphy/ BIA / Bald Eagle (Haliaeetus leucocephalus) fishing, Alaska

Alan Murphy/ BIA / Bald Eagle (Haliaeetus leucocephalus) fishing, Alaska

Markus Varesvuo / White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April.

Staffan Widstrand / Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January.

John Waters / Large murmuration of Common starlings (Sturnus vulgaris) swirling over reedbed roost at sunset, Ham Wall RSPB Reserve, Somerset, England, UK, January.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

Michael Hutchinson / Marsh fritillary butterfly (Euphydryas aurinia) flying and landing on a Buttercup (Ranunculus), Devon, England, UK, June.

Juan Carlos Munoz / Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March.

Pedro Narra / Bee hummingbird (Mellisuga helenae) hovering in front of flower, Guanahacabibes Peninsula National Park, Cuba

Pedro Narra / Barn Swallow (Hirundo rustica) in flight. Portugal, Europe

John Chan / Black skimmer (Rynchops niger) skimming water surface / feeding, Southern California, USA, July.

Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying over frost and snow covered forest. Tartumaa, Southern Estonia. January 2011.

Sven Zacek / Ural owl (Strix uralensis) female in flight having taken off from nest, in evening light. Tartumaa, Southern Estonia. May.

Sven Zacek / Ural owl (Strix uralensis) female flying with prey in beak. Flying back to nest in evening light. Tartumaa, Southern Estonia. May.

Melvin Grey / Purple-throated mountain gem (Lampornis calolaemus) adult female - about to feed from flower, Atlantic Rainforest, Costa Rica.

Paul Souders/ Worldfoto / Lesser Flamingo (Phoenicopterus minor) flock flying, Lake Magadi, Kenya

Paul Souders/ Worldfoto / Grey Crowned Crane (Balearica regulorum) flying, Ngorongoro Conservation Area, Tanzania

Konrad Wothe / Firefly (Lamprohiza splendidula) light tracks in forest, Bavaria, Germany

Konrad Wothe / Small Tortoiseshell (Aglais urticae) butterfly flying, Bavaria, Germany

Konrad Wothe / Honey Bee (Apis mellifera) flying with pollen basket, Bavaria, Germany

Ismaele Tortella / White tailed eagle (Haliaeetus albicilla) flying, Lofoten, Norway, August.

Morley Read / Slow motion clip of a large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Twistneck turtle (Platemys platycephala) resting, with butterfly landing on its head, Rio Yasuni, Orellana Province, Ecuador. (non-ex)