Tony Wu / Demon stinger (Inimicus didactylus) sitting on the sand with venomous spines on full display. Lembeh Strait, North Sulawesi.

Tony Wu / Humpback whales (Megaptera novaeangliae) engaged in bubble net feeding. Alaska, USA, July.

Tony Wu / Humpback whales (Megaptera novaeangliae kuzira) engaged in bubble-net feeding, highlighted by the warm light of a late summer evening. Kupreanof Island, Alaska, USA. July.

Tony Wu / Humpback whale (Megaptera novaeangliae) lying on its side while executing a series of pectoral fin slaps on ocean surface. Vava'u, Tonga, South Pacific.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female and calf swimming together at depth, ventral surfaces of both whales clearly visible. Vava'u, Tonga, Pacific Ocean.

Tony Wu / Water streaming off of a diving Humpback whale's (Megaptera novaeangliae) fluke, with 42ft Nordic Tug Legend visible in the background. Chantham Strait, Alaska, USA.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female breaching. Vava'u, Tonga. Pacific Ocean.

Tony Wu / Humpback whale calf (Megaptera novaeangliae) male calf peeking out from underneath his pectoral fin as he swims upside-down beneath the ocean surface. His mother was resting about 20 meters below. Vav...

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) female nudging her mother's mammary gland whilst trying to feed. Vava;u, Tonga, South Pacific.

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) male calf launching partially out of the water with his mouth open while playing together with mother. Vava'u, Tonga, Pacific Ocean.

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) swimming just under the ocean surface, partially backlit by dramatic rays of sunlight. Vava'u, Tonga, Pacific Ocean.

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis). faeces. Defecation like this from calves contributes to the cycling of nutrients from nutrient-rich high latitudes to less nutrient-abundant low...

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) female calf breaching sequence. Vava'u, Tonga, Pacific Ocean.

Tony Wu / North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July.

Tony Wu / North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching, Icy Strait, Alaska, USA. July.

Tony Wu / Developing eggs of Fat greenling (Hexagrammos otakii) Miyagi Prefecture, Japan. Pacific Ocean.

Tony Wu / Grunt sculpin (Rhamphocottus richardsoni) male watching over a clutch of eggs. Miyagi Prefecture, Japan, Pacific Ocean.

Tony Wu / Greater crested tern (Thalasseus bergii) feeding on Sardine, Tonga.

Alex Mustard / Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean...

Alex Mustard / Blacktip reef shark (Carcharhinus melanopterus) swims beneath a jetty and schools of silversides (Atherinidae) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Paci...

Alex Mustard / Few fish from a school of Bigeye trevally (+Caranx sexfasciatus) break off to scratch themselves against the flank of a blacktip reef shark (Carcharhinus melanopterus). This is a common and world wide...

Alex Mustard / Coral grouper (Cephalopholis miniata) waits in ambush, hiding against a seafan (Annella sp.) hunting silversides (Atherinidae), with schooling bigeye snapper (Lutjanus lutjanus) behind. Misool, Raja A...

Alex Mustard / Large reef scene on the current exposed face of a coral reef, with seafans (Melithaea sp.), soft corals (Dendronephthya spp.), sponges, coral grouper (Cephalopholis miniata), anthias, damselfish and m...

Alex Mustard / Commensal emperor shrimp (Periclimenes imperator) moves across the coloruful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...

Alex Mustard / Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips. Misool, Raja Ampat, West Papua, Indone...

Alex Mustard / Red sea fan (Melithaea sp.) growing beneath an island (Nudi Rock), with sun rays and trees. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island. Misool, Raja Ampat, West Papua, Indones...

Alex Mustard / Portrait of a large female Reef manta ray (Mobula alfredi) swimming over a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Long exposure of school of Bigeye trevally (Caranx sexfasciatus) swimming in front of a jetty, close to sunset. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Killer whale / Orca (Orcinus orca) newborn breaching to breath at the surface in an Arctic fjord in Spildra, northern Norway. This baby was so young that it could not yet breathe properly with its blo...

Alex Mustard / California sea lion (Zalophus californianus) pups in an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.

Alex Mustard / California sea lion pup (Zalophus californianus) practicings its hunting skills by playing chase with a falling sea star (Panamic cushion star: Pentacerster cumingi). Los Islotes, La Paz, Baja Califor...

Alex Mustard / Pair of California sea lion (Zalophus californianus) pups playing in shallow water. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.

Alex Mustard / Pair of crown-of-thorns sea stars (Acanthaster planci) feeding on a coral colony. La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.

Alex Mustard / Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind , in the plankton-rich green waters of La Paz bay, Baja...

Alex Mustard / Long exposure portrait of a Tawny nurse shark (Nebris ferrungineus), with Pink whipray (Himantura fai) and Bluefin trevally (Caranx melampygus) at dusk. Vavuu Atoll, Maldives. Indian Ocean

Alex Mustard / Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean

Alex Mustard / Several mimic goatfish (Mulliodichthys mimicus) hide within a school of Bluestripe snapper (Lutjanus kasmira) on a coral reef. North Ari Atoll, Maldives. Indian Ocean

Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean

Alex Mustard / Three moray eels, a Honeycomb moray: (Gymnothorax favagineus), Yellowmargin moray: (Gymnothorax flavimarginatus) and White-eyed moray (Gymnothorax thrysoideus) emerge from a single hole in a coral ree...

Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the packground sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...

Alex Mustard / Clark's anemonefish (Amphiprion clarkii) tends to its clutch of eggs. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Pygmy seahorse (Hippocampus bargibanti,) hiding in a seafan (Muricella sp.). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.Cropped.

Alex Mustard / Mushroom coral (Fungia sp.) fluoresces a bright orange at depth. At this depth underwater there is no orange light left, pigments within the coral absorb blue light and emit it as orange light (the re...

Alex Mustard / Mimic octopus (Thaumoctopus mimicus) swims over a sandy seabed in a shape considered by some to be a flounder impersonation. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Common seahorse (Hippocampus kuda). The dark patch beneath his snout is his brood pouch. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef. This photo shows the larger...

Alex Mustard / Mototi octopus (Amphioctopus mototi) swims across the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...

Alex Mustard / Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Veined octopus (Amphioctopus marginatus) shelters inside an old clam shell, with silver-sides in the water above. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Long exposure of a bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Blue ringed octopus (Hapalochlaena lunulata) moving across the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Group of five reef lizardifsh (Synodus variegatus) waiting in ambush. These were perched on the highest point of a coral reef attempting to catch schooling baitfish above. Bitung, North Sulawesi, Indo...

Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a colourful nudibranch (Chromodoris annae). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Staffan Widstrand / Rufous treepie (Dendrocitta vagabunda) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / White-breasted waterhen (Amaurornis phoenicurus) in Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Rose-ringed parakeet (Psittacula krameri) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Painted stork (Mycteria leucocephala) head and beak portrait, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian / Blue peafowl (Pavo cristatus) head portrait, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Grey francolin (Francolinus pondicerianus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Northern palm squirrel (Funambulus pennantii)Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian jungle crow (Corvus culminatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / White-breasted waterhen (Amaurornis phoenicurus) bathing, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Spotted dove (Spilopelia chinensis) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Grey francolin (Francolinus pondicerianus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Laughing dove (Spilopelia senegalensis) in Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India.

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Gaur or Indian bison (Bos gaurus / Bos frontalis) Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Barasingha / Swamp deer (Rucervus / Cervus duvaucelii) stag grooming, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Spotted deer or Chital (Axis axis) stag grooming its back, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Bengal tiger (Panthera tigris tigris) male resting in waterhole, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Gaur or Indian Bison (Bos gaurus), Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Bengal tiger (Panthera tigris tigris) male covered in mud, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Golden jackal (Canis aureus) Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Golden jackal (Canis aureus) with its kill, Spotted deer or Chital (Axis axis) fawn, in Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Golden jackal (Canis aureus) Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Indian flying fox or Greater Indian fruit bat (Pteropus giganteus) in Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Indian flying fox or Greater Indian fruit bat (Pteropus giganteus) in Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Staffan Widstrand / Great cormorant (Phalacrocorax carbo) in Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Large-tailed nightjar (Caprimulgus macrurus) in Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Iberian or Spanish imperial eagle (Aquila adalberti) eating a rabbit put out for it at a wildlife watching hide near El Barraco, Gredos Mountains, Avila, Spain.