Tom Vezo / Grey wolves showing fur colour variation. (Canis lupus)

Peter Blackwell / Lioness dragging zebra kill (Panthera leo) Masai Mara Africa

Anup Shah / Lioness with very young cub (Panthera leo) East Africa

Bernard Castelein / Lioness (Panthera leo) with cubs, Ngorongoro Crater, Tanzania

Barrie Britton / Juvenile Vervet monkey raids kitchen (Chlorocebus / Cercopithecus aethiops) Durban, South Africa Wildvision

Staffan Widstrand / Polar bears {Ursus maritimus} walking, Canada Wildvision Picture Publicity World's Best

DAVID TIPLING / Polish Tarpan horse (Equus caballus) grazing, Minsmere RSPB reserve, Suffolk, UK

WILLIAM OSBORN / Active nest of Median wasps {Dolichovespula media} hung in low bush UK