Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

Markus Varesvuo / Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.

Staffan Widstrand / Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January.

Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.

Sven Zacek / Ural owl (Strix uralensis) female stretching wings whilst perched in Spruce (Picea sp) tree. Tartumaa, Southern Estonia. May.

Sven Zacek / Ural owl (Strix uralensis) female in flight having taken off from nest, in evening light. Tartumaa, Southern Estonia. May.

Tui De Roy / Flightless cormorant (Phalacrocorax harrisi), two on rock. Bird drying wings in foreground with other sitting on nest in background. Cape Douglas, Fernandina Island, Galapagos.

Tui De Roy / Blue-footed booby (Sula nebouxii) landing. Punta Vicente Roca, Isabela Island, Galapagos.

Guy Edwardes / African fish eagle (Haliaeetus vocifer), Savuti, Botswana.

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick under the wing of protective parent, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June.Finalist in the Portfolio Category...

Melvin Grey / Violet-tailed sylph hummingbird (Aglaiocercus coelestis) hummingbird adult male, approaching flower to feed, Tandayapa, Ecuador