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Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS,  GDT 2011 competition
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Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition
Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
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Charlie Hamilton James / Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
Lesser flamingos {Phoeniconaias minor} performing 'wing salute' as part of courtship display, the wings are spread out to the side of the body and held in this position for several seconds, making the...
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Anup Shah / Lesser flamingos {Phoeniconaias minor} performing 'wing salute' as part of courtship display, the wings are spread out to the side of the body and held in this position for several seconds, making the...
Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 2 of 3
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Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 2 of 3
Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.
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Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.
Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
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Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April.
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Marie Read / Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April.
Baltimore Oriole (Icterus galbula) males fighting, Texas
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Alan Murphy/ BIA / Baltimore Oriole (Icterus galbula) males fighting, Texas
Baltimore Oriole (Icterus galbula) males fighting, Texas
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Alan Murphy/ BIA / Baltimore Oriole (Icterus galbula) males fighting, Texas
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.
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Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.
Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.
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Markus Varesvuo / Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
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George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.
Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
Black Heron (Egretta ardesiaca) canopy hunting, Chobe River, Chobe National Park, Botswana
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Paul Souders/ Worldfoto / Black Heron (Egretta ardesiaca) canopy hunting, Chobe River, Chobe National Park, Botswana
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.
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Paul Hobson / Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.
American Avocet (Recurvirostra americana) pair courting, Montana
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Donald M. Jones / American Avocet (Recurvirostra americana) pair courting, Montana
Sparkling Violet-ear (Colibri coruscans) hummingbird bathing, Bogota, Colombia
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Thomas Marent / Sparkling Violet-ear (Colibri coruscans) hummingbird bathing, Bogota, Colombia
Common Loon (Gavia immer) spreading wings, Monterey Bay, California
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Chase Dekker / Common Loon (Gavia immer) spreading wings, Monterey Bay, California
Northern Cardinal (Cardinalis cardinalis) male and Pyrrhuloxia (Cardinalis sinuatus) male fighting, Texas
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Alan Murphy/ BIA / Northern Cardinal (Cardinalis cardinalis) male and Pyrrhuloxia (Cardinalis sinuatus) male fighting, Texas
Gentoo Penguin (Pygoscelis papua) wading, Antarctica
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Benjamin Olson / Gentoo Penguin (Pygoscelis papua) wading, Antarctica
Gentoo Penguin (Pygoscelis papua) wet from rain due to climate change, Antarctica
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Benjamin Olson / Gentoo Penguin (Pygoscelis papua) wet from rain due to climate change, Antarctica
Common Loon (Gavia immer) stretching, Troy, Montana
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Donald M. Jones / Common Loon (Gavia immer) stretching, Troy, Montana
Grey Heron (Ardea cinerea) spreading wings on frozen lake, Berlin, Germany
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Jan Wegener/ BIA / Grey Heron (Ardea cinerea) spreading wings on frozen lake, Berlin, Germany
Flightless cormorant (Phalacrocorax harrisi) silhouetted,  Punta Espinosa, Fernandina Island, Galapagos
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Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) silhouetted, Punta Espinosa, Fernandina Island, Galapagos
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June.
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Guy Edwardes / Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June.
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016.
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Ross Hoddinott / Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016.
Brown Pelican (Pelecanus occidentalis) hunting baitfish, Urvina Bay, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Brown Pelican (Pelecanus occidentalis) hunting baitfish, Urvina Bay, Isabela Island, Galapagos Islands, Ecuador
Fulgorid Planthopper (Phrictus quinquepartitus) in defensive posture, Braulio Carrillo National Park, Costa Rica
00572135
Paul Bertner / Fulgorid Planthopper (Phrictus quinquepartitus) in defensive posture, Braulio Carrillo National Park, Costa Rica
Pair of Blue footed boobies (Sula nebouxii) courting, with the male picking up and offering grass, Isla Lobos de Terra, Peru.
01534923
Fred Olivier / Pair of Blue footed boobies (Sula nebouxii) courting, with the male picking up and offering grass, Isla Lobos de Terra, Peru.
Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. June 2014
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David Pattyn / Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. June 2014
Baya weaver (Ploceus philippinus) male flying to nest entrance, Singapore
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Ingo Arndt / Baya weaver (Ploceus philippinus) male flying to nest entrance, Singapore
Kingfisher (Alcedo atthis) leaving water after successfully catching a fish, Hessen, Germany, sequence 9/9
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Ingo Arndt / Kingfisher (Alcedo atthis) leaving water after successfully catching a fish, Hessen, Germany, sequence 9/9
Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 7/9
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Ingo Arndt / Kingfisher (Alcedo atthis) diving into water to catch a fish, Hessen, Germany, sequence 7/9
Purple Heron (Ardea purpurea) pair courting and building nest in reeds, Germany
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Ingo Arndt / Purple Heron (Ardea purpurea) pair courting and building nest in reeds, Germany
Velvet-purple Coronet (Boissonneaua jardini) hummingbird in defensive posture, Las Tangaras Bird Reserve, Colombia
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Thomas Marent / Velvet-purple Coronet (Boissonneaua jardini) hummingbird in defensive posture, Las Tangaras Bird Reserve, Colombia
Pied Kingfisher (Ceryle rudis) female landing, Gambia
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Ralf Kistowski/ BIA / Pied Kingfisher (Ceryle rudis) female landing, Gambia
Golden-breasted Puffleg (Eriocnemis mosquera) feeding on flower nectar, Colombia
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Glenn Bartley/ BIA / Golden-breasted Puffleg (Eriocnemis mosquera) feeding on flower nectar, Colombia
Beautiful Demoiselle (Calopteryx virgo), North Rhine-Westphalia, Germany
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Ralf Kistowski/ BIA / Beautiful Demoiselle (Calopteryx virgo), North Rhine-Westphalia, Germany
Dryad (Minois dryas) butterfly, Poland
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Peter Schwarz/ BIA / Dryad (Minois dryas) butterfly, Poland
Common Kingfisher (Alcedo atthis) parent spreading wings near fledgling, North Rhine-Westphalia, Germany
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Ralf Kistowski/ BIA / Common Kingfisher (Alcedo atthis) parent spreading wings near fledgling, North Rhine-Westphalia, Germany
Black-winged Lapwing (Vanellus melanopterus) calling during defensive display, KwaZulu-Natal, South Africa
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Eric Sohn Joo Tan/ BIA / Black-winged Lapwing (Vanellus melanopterus) calling during defensive display, KwaZulu-Natal, South Africa
Montagu's Harrier (Circus pygargus) male landing, Andalusia, Spain
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Alain Ghignone/ BIA / Montagu's Harrier (Circus pygargus) male landing, Andalusia, Spain
Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany
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Thomas Hinsche/ BIA / Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany
Eurasian Hoopoe (Upupa epops) stretching, Aosta Valley, Italy
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Alain Ghignone/ BIA / Eurasian Hoopoe (Upupa epops) stretching, Aosta Valley, Italy
Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany
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Thomas Hinsche/ BIA / Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany
Say's Phoebe (Sayornis saya) landing, Texas
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Alan Murphy/ BIA / Say's Phoebe (Sayornis saya) landing, Texas
Griffon vulture (Gyps fulvus) coming in to land, Spain, July
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David Kjaer / Griffon vulture (Gyps fulvus) coming in to land, Spain, July
Bearded vulture (Gypaetus barbatus)) coming in to land, Spain, July
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David Kjaer / Bearded vulture (Gypaetus barbatus)) coming in to land, Spain, July
Egytian Goose (Alopochen aegyptiaca) rear view  bathing, sings spread, Zimanga Private Game Reserves, South Africa
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Wim van den Heever / Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa
Grey herons (Ardea cinerea) fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
01531815
Bence Mate / Grey herons (Ardea cinerea) fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. June.
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Bence Mate / Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. June.
Piping Plover (Charadrius melodus) stretching, Massachusetts
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Michael Milicia/ BIA / Piping Plover (Charadrius melodus) stretching, Massachusetts
Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February
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David Pattyn / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February
Garden warbler (Sylvia borin), catching flies in willow tree, Korpoo, Uto, Finland, May.
01526632
Jussi Murtosaari / Garden warbler (Sylvia borin), catching flies in willow tree, Korpoo, Uto, Finland, May.
Grey heron (Ardea cinerea)  landing on lake, Pusztaszer, Hungary, April
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Michel Poinsignon / Grey heron (Ardea cinerea) landing on lake, Pusztaszer, Hungary, April
Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.
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Wim van den Heever / Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.
White-fronted bee-eaters (Merops bullockoides) perched near to nesting holes Chobe National Park, Botswana.
01529771
Wim van den Heever / White-fronted bee-eaters (Merops bullockoides) perched near to nesting holes Chobe National Park, Botswana.
Black heron (Egretta ardesiaca) fishing and using wings to create an area of shade to attract fish, on the banks of the Chobe River, Botswana.
01529760
Wim van den Heever / Black heron (Egretta ardesiaca) fishing and using wings to create an area of shade to attract fish, on the banks of the Chobe River, Botswana.
White tailed eagle (Haliaeetus albicilla) caught in a fight. Hokkaido Japan, March.
01529747
Wim van den Heever / White tailed eagle (Haliaeetus albicilla) caught in a fight. Hokkaido Japan, March.
White tailed sea eagle (Haliaeetus albicilla) flying looking over shoulder, Hokkaido Japan.
01529743
Wim van den Heever / White tailed sea eagle (Haliaeetus albicilla) flying looking over shoulder, Hokkaido Japan.
Japanese crane (Grus japonensis) jumping in the air, Hokkaido, Japan, March.
01529742
Wim van den Heever / Japanese crane (Grus japonensis) jumping in the air, Hokkaido, Japan, March.
Whooper swan (Cygnus cygnus) mid flight over the frozen lake, Kussharo, Hokkaido, Japan.
01529735
Wim van den Heever / Whooper swan (Cygnus cygnus) mid flight over the frozen lake, Kussharo, Hokkaido, Japan.
Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.
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Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.
Australasian gannet (Morus serrator) flying with waves breaking below, Muriwai, Auckland, New Zealand, October.
01527557
Brent Stephenson / Australasian gannet (Morus serrator) flying with waves breaking below, Muriwai, Auckland, New Zealand, October.
Wood Duck (Aix sponsa) drake stretching wings, central Montana
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Donald M. Jones / Wood Duck (Aix sponsa) drake stretching wings, central Montana
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.
01520518
Wim van den Heever / Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.
Ruff (Philomachus pugnax) male displaying at lek, Varanger Peninsula, Norway
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Winfried Wisniewski / Ruff (Philomachus pugnax) male displaying at lek, Varanger Peninsula, Norway
Snowy Egret (Egretta thula) four week old chick begging for food, Sonoma County, California
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Suzi Eszterhas / Snowy Egret (Egretta thula) four week old chick begging for food, Sonoma County, California
Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.
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Sven Zacek / Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.
Four spotted chaser (Libellula quardrimaculata) Montiaghs Moss NNR, Aghalee, County Antrim, Northern Ireland, June.
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Robert Thompson / Four spotted chaser (Libellula quardrimaculata) Montiaghs Moss NNR, Aghalee, County Antrim, Northern Ireland, June.
Tolga Bat Hospital volunteer handing Little red flying fox (Pteropus scapulatus) baby as it flaps her wings, Tolga Bat Hospital, Atherton Tablelands, Queensland, Australia. May 2015. Second Place in t...
01520166
Juergen Freund / Tolga Bat Hospital volunteer handing Little red flying fox (Pteropus scapulatus) baby as it flaps her wings, Tolga Bat Hospital, Atherton Tablelands, Queensland, Australia. May 2015. Second Place in t...
Osprey (Pandion haliaetus) in flight with nesting material, Hungary, July. Nominated in the Telephoto category of the Terre Sauvage Nature Images Awards competition 2015.
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Bence Mate / Osprey (Pandion haliaetus) in flight with nesting material, Hungary, July. Nominated in the Telephoto category of the Terre Sauvage Nature Images Awards competition 2015.
Black-throated diver (Gavia arctica) displaying, Finland, June.
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Danny Green / Black-throated diver (Gavia arctica) displaying, Finland, June.
Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa.
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Neil Aldridge / Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa.
Adelie Penguin (Pygoscelis adeliae) carrying stolen rock from another nest in rookery, Peterman Island, Antarctica, November.
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Charlie Summers / Adelie Penguin (Pygoscelis adeliae) carrying stolen rock from another nest in rookery, Peterman Island, Antarctica, November.
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July.
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Marie Read / Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July.
White-tailed sea eagle (Haliaeetus albicilla) in flight, against a clear blue sky, Norway, July.
01515781
Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) in flight, against a clear blue sky, Norway, July.
White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.
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Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.
White-tailed sea eagle (Haliaeetus albicilla) flying, against a clear blue sky, Norway, July.
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Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, against a clear blue sky, Norway, July.
Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
Fulmar (Fulmaris glacialis) in flight with Hunstanton, Norfolk, England, UK, February.
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Ernie Janes / Fulmar (Fulmaris glacialis) in flight with Hunstanton, Norfolk, England, UK, February.
Gyrfalcon (Falco rusticolus) and Raven (Corvus corax) in flight, Hornoya bird cliff, Finnmark, Norway. March
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Espen Bergersen / Gyrfalcon (Falco rusticolus) and Raven (Corvus corax) in flight, Hornoya bird cliff, Finnmark, Norway. March
Gyrfalcon (Falco rusticolus) in flight with guillemot prey. Hornoya birdcliff. Finnmark, Norway. March
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Espen Bergersen / Gyrfalcon (Falco rusticolus) in flight with guillemot prey. Hornoya birdcliff. Finnmark, Norway. March
Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March
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Espen Bergersen / Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March
Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.
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Espen Bergersen / Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
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Danny Green / Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
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Danny Green / Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
Steller's sea eagle (Haliaeetus pelagicus) in flight, Japan, February
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Danny Green / Steller's sea eagle (Haliaeetus pelagicus) in flight, Japan, February
Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February
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Danny Green / Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February
Emperor moth(Saturnia pavonia) female, Peak District National Park, UK. May.
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Alex Hyde / Emperor moth(Saturnia pavonia) female, Peak District National Park, UK. May.
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April.
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Alex Hyde / Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April.
Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
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Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
Nazca booby (Sula granti) in flight with rainbow behind, Galapagos.
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Ole Jorgen Liodden / Nazca booby (Sula granti) in flight with rainbow behind, Galapagos.
Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
01507140
Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.