Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition

Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

Markus Varesvuo / Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.

George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.

Paul Hobson / Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.

Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) silhouetted, Punta Espinosa, Fernandina Island, Galapagos

Ross Hoddinott / Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016.

Frederique Olivier / Pair of Blue footed boobies (Sula nebouxii) courting, with the male picking up and offering grass, Isla Lobos de Terra, Peru.

David Pattyn / Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. June 2014