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Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS,  GDT 2011 competition
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Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition
Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
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Charlie Hamilton James / Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.
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Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.
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Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.
Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.
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Markus Varesvuo / Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
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George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.
Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.
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Paul Hobson / Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.
Flightless cormorant (Phalacrocorax harrisi) silhouetted,  Punta Espinosa, Fernandina Island, Galapagos
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Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) silhouetted, Punta Espinosa, Fernandina Island, Galapagos
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016.
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Ross Hoddinott / Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016.
Pair of Blue footed boobies (Sula nebouxii) courting, with the male picking up and offering grass, Isla Lobos de Terra, Peru.
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Fred Olivier / Pair of Blue footed boobies (Sula nebouxii) courting, with the male picking up and offering grass, Isla Lobos de Terra, Peru.
Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. June 2014
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David Pattyn / Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. June 2014
Baya weaver (Ploceus philippinus) male flying to nest entrance, Singapore
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Ingo Arndt / Baya weaver (Ploceus philippinus) male flying to nest entrance, Singapore
Griffon vulture (Gyps fulvus) coming in to land, Spain, July
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David Kjaer / Griffon vulture (Gyps fulvus) coming in to land, Spain, July
Bearded vulture (Gypaetus barbatus)) coming in to land, Spain, July
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David Kjaer / Bearded vulture (Gypaetus barbatus)) coming in to land, Spain, July
Grey herons (Ardea cinerea) fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
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Bence Mate / Grey herons (Ardea cinerea) fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. June.
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Bence Mate / Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. June.
Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February
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David Pattyn / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February
Garden warbler (Sylvia borin), catching flies in willow tree, Korpoo, Uto, Finland, May.
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Jussi Murtosaari / Garden warbler (Sylvia borin), catching flies in willow tree, Korpoo, Uto, Finland, May.
Grey heron (Ardea cinerea)  landing on lake, Pusztaszer, Hungary, April
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Michel Poinsignon / Grey heron (Ardea cinerea) landing on lake, Pusztaszer, Hungary, April
Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.
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Wim van den Heever / Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.
White-fronted bee-eaters (Merops bullockoides) perched near to nesting holes Chobe National Park, Botswana.
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Wim van den Heever / White-fronted bee-eaters (Merops bullockoides) perched near to nesting holes Chobe National Park, Botswana.
Black heron (Egretta ardesiaca) fishing and using wings to create an area of shade to attract fish, on the banks of the Chobe River, Botswana.
01529760
Wim van den Heever / Black heron (Egretta ardesiaca) fishing and using wings to create an area of shade to attract fish, on the banks of the Chobe River, Botswana.
White tailed eagle (Haliaeetus albicilla) caught in a fight. Hokkaido Japan, March.
01529747
Wim van den Heever / White tailed eagle (Haliaeetus albicilla) caught in a fight. Hokkaido Japan, March.
White tailed sea eagle (Haliaeetus albicilla) flying looking over shoulder, Hokkaido Japan.
01529743
Wim van den Heever / White tailed sea eagle (Haliaeetus albicilla) flying looking over shoulder, Hokkaido Japan.
Japanese crane (Grus japonensis) jumping in the air, Hokkaido, Japan, March.
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Wim van den Heever / Japanese crane (Grus japonensis) jumping in the air, Hokkaido, Japan, March.
Whooper swan (Cygnus cygnus) mid flight over the frozen lake, Kussharo, Hokkaido, Japan.
01529735
Wim van den Heever / Whooper swan (Cygnus cygnus) mid flight over the frozen lake, Kussharo, Hokkaido, Japan.
Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.
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Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.
Australasian gannet (Morus serrator) flying with waves breaking below, Muriwai, Auckland, New Zealand, October.
01527557
Brent Stephenson / Australasian gannet (Morus serrator) flying with waves breaking below, Muriwai, Auckland, New Zealand, October.
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.
01520518
Wim van den Heever / Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.
Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.
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Sven Zacek / Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.
Four spotted chaser (Libellula quardrimaculata) Montiaghs Moss NNR, Aghalee, County Antrim, Northern Ireland, June.
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Robert Thompson / Four spotted chaser (Libellula quardrimaculata) Montiaghs Moss NNR, Aghalee, County Antrim, Northern Ireland, June.
Tolga Bat Hospital volunteer handing Little red flying fox (Pteropus scapulatus) baby as it flaps her wings, Tolga Bat Hospital, Atherton Tablelands, Queensland, Australia. May 2015. Second Place in t...
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Juergen Freund / Tolga Bat Hospital volunteer handing Little red flying fox (Pteropus scapulatus) baby as it flaps her wings, Tolga Bat Hospital, Atherton Tablelands, Queensland, Australia. May 2015. Second Place in t...
Osprey (Pandion haliaetus) in flight with nesting material, Hungary, July. Nominated in the Telephoto category of the Terre Sauvage Nature Images Awards competition 2015.
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Bence Mate / Osprey (Pandion haliaetus) in flight with nesting material, Hungary, July. Nominated in the Telephoto category of the Terre Sauvage Nature Images Awards competition 2015.
Black-throated diver (Gavia arctica) displaying, Finland, June.
01520897
Danny Green / Black-throated diver (Gavia arctica) displaying, Finland, June.
Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa.
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Neil Aldridge / Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa.
Adelie Penguin (Pygoscelis adeliae) carrying stolen rock from another nest in rookery, Peterman Island, Antarctica, November.
01514836
Charlie Summers / Adelie Penguin (Pygoscelis adeliae) carrying stolen rock from another nest in rookery, Peterman Island, Antarctica, November.
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July.
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Marie Read / Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July.
White-tailed sea eagle (Haliaeetus albicilla) in flight, against a clear blue sky, Norway, July.
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Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) in flight, against a clear blue sky, Norway, July.
White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.
01515779
Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.
White-tailed sea eagle (Haliaeetus albicilla) flying, against a clear blue sky, Norway, July.
01515776
Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, against a clear blue sky, Norway, July.
Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
01514090
Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
Fulmar (Fulmaris glacialis) in flight with Hunstanton, Norfolk, England, UK, February.
01512588
Ernie Janes / Fulmar (Fulmaris glacialis) in flight with Hunstanton, Norfolk, England, UK, February.
Gyrfalcon (Falco rusticolus) and Raven (Corvus corax) in flight, Hornoya bird cliff, Finnmark, Norway. March
01510329
Espen Bergersen / Gyrfalcon (Falco rusticolus) and Raven (Corvus corax) in flight, Hornoya bird cliff, Finnmark, Norway. March
Gyrfalcon (Falco rusticolus) in flight with guillemot prey. Hornoya birdcliff. Finnmark, Norway. March
01510328
Espen Bergersen / Gyrfalcon (Falco rusticolus) in flight with guillemot prey. Hornoya birdcliff. Finnmark, Norway. March
Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March
01510326
Espen Bergersen / Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March
Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.
01510350
Espen Bergersen / Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
01510232
Danny Green / Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
01510231
Danny Green / Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
Steller's sea eagle (Haliaeetus pelagicus) in flight, Japan, February
01510219
Danny Green / Steller's sea eagle (Haliaeetus pelagicus) in flight, Japan, February
Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February
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Danny Green / Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February
Emperor moth(Saturnia pavonia) female, Peak District National Park, UK. May.
01510033
Alex Hyde / Emperor moth(Saturnia pavonia) female, Peak District National Park, UK. May.
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April.
01509989
Alex Hyde / Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April.
Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
01507141
Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
Nazca booby (Sula granti) in flight with rainbow behind, Galapagos.
01507323
Ole Jorgen Liodden / Nazca booby (Sula granti) in flight with rainbow behind, Galapagos.
Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
01507140
Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
Light-mantled sooty albatross (Phoebetria palpebrata) in flight, Auckland Island, Sub-Antarctic New Zealand. February.
01507151
Ole Jorgen Liodden / Light-mantled sooty albatross (Phoebetria palpebrata) in flight, Auckland Island, Sub-Antarctic New Zealand. February.
Great grey owl (Strix nebulosa lapponica) in flight, close up, Finland, March.
01507181
Ole Jorgen Liodden / Great grey owl (Strix nebulosa lapponica) in flight, close up, Finland, March.
Great grey owl (Strix nebulosa lapponica) hunting in flight, Lapland, Finland, March.
01507276
Ole Jorgen Liodden / Great grey owl (Strix nebulosa lapponica) hunting in flight, Lapland, Finland, March.
Arctic tern (Sterna paridisea) mobbing, Arctic fox (Alopex lagopus) in Spitsbergen, Svalbard, Norway, July.
01507213
Ole Jorgen Liodden / Arctic tern (Sterna paridisea) mobbing, Arctic fox (Alopex lagopus) in Spitsbergen, Svalbard, Norway, July.
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species.
01506364
David Pattyn / Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species.
Shag (Phalacrocorax aristotelis) spreading wings to dry, Hornoya, Varanger, Finnmark, Norway, April.
01498232
David Pattyn / Shag (Phalacrocorax aristotelis) spreading wings to dry, Hornoya, Varanger, Finnmark, Norway, April.
Eagle hunter mounted on Mongolian horse with his female Golden eagle (Aquila chrysaetos) at Eagle Hunters Festival, near Sagsai, Bayan-Ulgii Aymag, Mongolia. September 2014..
01501633
Kristel Richard / Eagle hunter mounted on Mongolian horse with his female Golden eagle (Aquila chrysaetos) at Eagle Hunters Festival, near Sagsai, Bayan-Ulgii Aymag, Mongolia. September 2014..
Black kite (Milvus migrans) flying and red-crowned crane (Grus japonensis) on snow, Hokkaido, Japan, February.
01498044
Markus Varesvuo / Black kite (Milvus migrans) flying and red-crowned crane (Grus japonensis) on snow, Hokkaido, Japan, February.
Green-veined white butterfly (Pieris napi) in flight, Jyvaskyla, Keski-Finland / Central Finland, Finland, September.
01500799
Jussi Murtosaari / Green-veined white butterfly (Pieris napi) in flight, Jyvaskyla, Keski-Finland / Central Finland, Finland, September.
Apollo butterfly (Parnassius apollo) in flight, Foglo, Ahvenanmaa / Aland Islands, Finland, July.
01500789
Jussi Murtosaari / Apollo butterfly (Parnassius apollo) in flight, Foglo, Ahvenanmaa / Aland Islands, Finland, July.
Scarce tortoiseshell butterfly (Nymphalis xanthomelas) on pussy willow, Luhanka, Keski-Finland, April. / Central Finland, Finland, April.
01500774
Jussi Murtosaari / Scarce tortoiseshell butterfly (Nymphalis xanthomelas) on pussy willow, Luhanka, Keski-Finland, April. / Central Finland, Finland, April.
Arctic tern (Sterna Paradisaea) flying, Svalbard, Norway, July.
01501139
Ole Jorgen Liodden / Arctic tern (Sterna Paradisaea) flying, Svalbard, Norway, July.
Barn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird.
01492348
John Waters / Barn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird.
White-vented storm petrel (Oceanites gracilis galapagoensis) 'walking' on surface of water whilst foraging, Galapagos
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Tui De Roy / White-vented storm petrel (Oceanites gracilis galapagoensis) 'walking' on surface of water whilst foraging, Galapagos
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species.
01497290
Tui De Roy / Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species.
Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. April.
01492187
David Pattyn / Great crested grebe (Podiceps cristatus) stretching its wings. The Netherlands. April.
Great crested grebe (Podiceps cristatus) adult running over the water surface during the courtship. The Netherlands. April 2014.Second place in the Species Story category of the Terre Sauvage Nature I...
01492143
David Pattyn / Great crested grebe (Podiceps cristatus) adult running over the water surface during the courtship. The Netherlands. April 2014.Second place in the Species Story category of the Terre Sauvage Nature I...
Male King eiders (Somateria spectabilis) three taking-off from water. Batsfjord, Norway, March.
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Espen Bergersen / Male King eiders (Somateria spectabilis) three taking-off from water. Batsfjord, Norway, March.
Black guillemot (Cepphus grylle) taking off, Denmark, May
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Hanne & Jens Eriksen / Black guillemot (Cepphus grylle) taking off, Denmark, May
White-tailed Eagle (Haliaeetus albicilla) in flight with mountains in background, Hokkaido, Japan, February
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Markus Varesvuo / White-tailed Eagle (Haliaeetus albicilla) in flight with mountains in background, Hokkaido, Japan, February
Steller's Eagle (Haliaeteus pelagicus) in flight over snow, Hokkaido, Japan, February
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Markus Varesvuo / Steller's Eagle (Haliaeteus pelagicus) in flight over snow, Hokkaido, Japan, February
Steller's Eagle (Haliaeteus pelagicus) flying, with black background, Hokkaido, Japan, February
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Markus Varesvuo / Steller's Eagle (Haliaeteus pelagicus) flying, with black background, Hokkaido, Japan, February
Snow geese (Anser caerulescens) flying toward their gathering pond, with moon in the background,  Boque del Apache National Wildlife Refuge, New Mexico. December.
01489727
Jack Dykinga / Snow geese (Anser caerulescens) flying toward their gathering pond, with moon in the background, Boque del Apache National Wildlife Refuge, New Mexico. December.
Male Andean Condor (Vultur gryphus) flying amongst the clouds, Los Cuernos Peaks, Torres del Paine, Chile. March.
01489707
Jack Dykinga / Male Andean Condor (Vultur gryphus) flying amongst the clouds, Los Cuernos Peaks, Torres del Paine, Chile. March.
European Herring Gull (Larus argentatus) landing on the waters of a fjord,  Norway, May.
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Ben Cranke / European Herring Gull (Larus argentatus) landing on the waters of a fjord, Norway, May.
Golden Eagle (Aquila chrysaetos) in flight, Finland.  February.
01479645
Ben Cranke / Golden Eagle (Aquila chrysaetos) in flight, Finland. February.
Steller's Sea Eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan.  February.
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Ben Cranke / Steller's Sea Eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan. February.
Peregrine falcons (Falco peregrinus), juvenile female and male in flight, fighting over prey. Bristol, UK. June.
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Sam Hobson / Peregrine falcons (Falco peregrinus), juvenile female and male in flight, fighting over prey. Bristol, UK. June.
Peregrine falcon (Falco peregrinus), juvenile in flight with remains of feral pigeon. Bristol, UK. July.
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Sam Hobson / Peregrine falcon (Falco peregrinus), juvenile in flight with remains of feral pigeon. Bristol, UK. July.
Brown pelican (Pelecanus occidentalis) diving for fish with the Athala II boat in the background, Espumilla beach, Galapagos, Ecuador. April.
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Ross Hoddinott / Brown pelican (Pelecanus occidentalis) diving for fish with the Athala II boat in the background, Espumilla beach, Galapagos, Ecuador. April.
Northern goshawk (Accipiter gentilis), young adult male taking flight from building with prey. Berlin, Germany. March. Nominated in the Melvita Nature Images Awards competition 2014.
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Sam Hobson / Northern goshawk (Accipiter gentilis), young adult male taking flight from building with prey. Berlin, Germany. March. Nominated in the Melvita Nature Images Awards competition 2014.
Gyrfalcon (Falco rusticolus) in flight, from directly below, Norway, July. Magic Moments book plate.
01287672
Markus Varesvuo / Gyrfalcon (Falco rusticolus) in flight, from directly below, Norway, July. Magic Moments book plate.
Great Grey Owl (Strix nebulosa) hunting, Tornio, Finland, March
01287657
Markus Varesvuo / Great Grey Owl (Strix nebulosa) hunting, Tornio, Finland, March
Rough-legged Buzard (Buteo lagopus)  flying over water, Porvoo Finland
01261661
Markus Varesvuo / Rough-legged Buzard (Buteo lagopus) flying over water, Porvoo Finland
Kittiwake (Rissa tridactylus) flying through snow, Norway, April.
01243968
Markus Varesvuo / Kittiwake (Rissa tridactylus) flying through snow, Norway, April.
Bar-tailed godwit (Limosa lapponica) in flight over water, Hanko, Finland, September
01261398
Markus Varesvuo / Bar-tailed godwit (Limosa lapponica) in flight over water, Hanko, Finland, September
Common chaffinchs (Fringilla coelebs) fighting, Puszaszer, Hungary, May.
01464869
Bence Mate / Common chaffinchs (Fringilla coelebs) fighting, Puszaszer, Hungary, May.
Red and green macaws (Ara chloropterus) in flight, motion blurred photograph, Buraxo das aras, Brazil.
01464865
Bence Mate / Red and green macaws (Ara chloropterus) in flight, motion blurred photograph, Buraxo das aras, Brazil.