Nature Production / Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.

Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition

Nick Gordon / Silky marmosets {Callithrix humeralifer chrysoleuca} aggression, Amazonia, Brazil

Anup Shah / African fish eagle {Haliaeetus vocifer} defends Lesser flamingo carcass from Marabou storks {Leptoptilos crumeniferus}, Lake Nakuru, Kenya.

Anup Shah / Juvenile Tawny or African Sea Eagle defending flamingo carcass from Marabou stork {Lepitoptilos crumeniferous}

Anup Shah / African fish eagle {Haliaeetus vocifer} fighting Marabou storks {Leptoptilos crumeniferus} over Lesser flamingo carcass, Lake Nakuru, Kenya.

Doug Perrine / Great white shark jaws open at surface {Carcharodon carcharias} South Africa (Non-ex).

Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa

Staffan Widstrand / Polar Bear (Ursus maritimus) mobbed by possibly a Skua, Svalbard, Norway, July

Paul Hobson / Arctic tern (Sterna paradisaea) two fighting in the air, Scotland, UK July

Denis-Huot / Masai Giraffe (Giraffa camelopardalis tippelskirchi), males fighting. Masai-Mara game reserve, Kenya.

Andrew Walmsley / Arctic terns (Sterna paradisaea), adults defending their nests from photographers, Northumberland, June. No release available.

Igor Shpilenok / Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 4...

Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March

David Pattyn / Siskins (Carduelis spinus) fighting over food. Oisterwijk, The Netherlands, March.

Juan Carlos Munoz / Bearded Vultures (Gypaetus barbatus) juvenile (left) and adult fighting in flight. Simien National Park, Ethiopia, Africa.

Mark Carwardine / Amazon river dolphin, pink river dolphin or boto (Inia geoffrensis) Rio Negro, Brazil (Amazon) - two wild animals fighting over fish Threatened species (IUCN Red List)

Mark Carwardine / Humpback whale lobtailing {Megaptera novaengliae} Dominican Republic, Caribbean

Mark Carwardine / Humpback whale lobtailing {Megaptera novaengliae} Dominican Republic, Caribbean

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / Royal Penguins (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Christophe Courteau / Fighting between a pack of 21 African Wild Dogs {Lycaon pictus} and the dominant female of a clan of Spotted Hyaenas (Crocuta crocuta) Northern Okavango Delta, Botswana

Christophe Courteau / Fighting between a pack of 21 African Wild Dogs {Lycaon pictus} and the dominant female of a clan of Spotted Hyaenas (Crocuta crocuta) Northern Okavango Delta, Botswana

Christophe Courteau / Two African Buffalo {Sincerus caffer} bulls sparring, Okavango Delta, Botswana.

Stefan Huwiler / Golden Eagle (Aquila chrysaetos), adult defending carcass from Red Fox (Vulpes vulpes), Sinite Kamani National Park, Bulgaria, Europe. February

Stefan Huwiler / Golden Eagle (Aquila chrysaetos) adult defending carcass from Red Fox (Vulpes vulpes), Sinite Kamani National Park, Bulgaria, Europe. February. Commended, Behaviour : Birds category, Wildlife Photogra...

Andy Trowbridge / Yellow-eyed Penguins (Megadyptes antipodes) vocalising with chick. Endangered species. Katiki Point, Moeraki, Otago, New Zealand. January.

Anup Shah / Mature male gelada (Theropithecus gelada) fighting head-on, upper lip curled back revealing teeth. Simien Mountains National Park, Ethiopia.

Kim Taylor / Wood Ant (Formica rufa) worker protecting the nest by jetting formic acid from its tail. Endemic to Europe and Asia. Surrey, UK.

David Pattyn / Red foxes (Vulpes vulpes) two fighting, probably over territory, The Netherlands, August

David Pattyn / Red necked grebe (Podiceps grisegena) approaching a competitor in an attempt to drive him away during the courting season, Danube area, Bulgaria, April

Juan Carlos Munoz / Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Great Egrets (Ardea alba) territorial dispute above nest colony, Everglades National Park, Florida, USA, February.

Martin Dohrn / Maricopa harvester ant (Pogonomyrmex maricopa) fighting a different species of ant (Aphaenogaster cockerelli), over territory Arizona, USA

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male charging at an African lion (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 9/11

Andrew Parkinson / Redwing (Turdus iliacus) protecting a windfall apple from another bird. Derbyshire, UK, February. (Non-ex)

Andrew Parkinson / Redwings (Turdus iliacus) squabbling over an apple in snow. Derbyshire, UK, February. British Wildlife Photographer of the Year (BWPA) competition 2012, 'Animal Behaviour' category. (Non-ex) Winner of...

Andy Rouse / European hares (Lepus europaeus) boxing, UK

Andy Rouse / European hares (Lepus europaeus) boxing, UK

Mark Hamblin / 2020VISION / Two Goldfinches (Carduelis carduelis) squabbling over Common teasel (Dipsacus fullonum) seeds in winter, Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. 2020VISION Exhibition.

Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank whilst third looks on from above (Panthera onca) from boat on river, Pantanal, Pocone, Brazil. No release available.

Andy Rouse / 2020VISION / European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 2 of 2. Did you know? A group of hares is known as a drove.

Andy Rouse / 2020VISION / European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2.

Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia

Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia

Charlie Summers / Plains Zebra (Equus quagga) in a dominance fight with dust flying, Lake Magadi, Serengeti, Tanzania.

Edwin Giesbers / Alpine marmots (Marmota marmota) fighting, Hohe Tauern National Park, Austria, July

Carol Walker / Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010

Kim Taylor / Two Siskins (Carduelis spinus) fighting, Surrey, UK, March

Mark Hamblin / 2020VISION / Goldfinches (Carduelis carduelis) squabbling over teasel seeds in winter. Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. Crop of 01390677

Mark Hamblin / 2020VISION / Mountain Hares (Lepus timidus) boxing in winter. Cairngorms National Park, Scotland, UK, January. Did you know? Although they are normally solitary, Mountain hares can gather in groups of up to 20 dur...

Danny Green / 2020VISION / Two European river otters (Lutra lutra) fighting over a fish, Isle of Mull, Inner Hebrides, Scotland, UK, December

Danny Green / 2020VISION / European river otters (Lutra lutra) play fighting amongst seaweed, Isle of Mull, Inner Hebrides, Scotland, UK, December

Colin Varndell / Greenfinches (Carduelis chloris) squabbling in flight, Dorset, UK, February

George Sanker / Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA

Daniel Heuclin / Green tree snake (Phyllodryas viridissima) on a branch about to stike, Bolivian Amazonia, controlled conditions

Orsolya Haarberg / Two Black grouse (Tetrao tetrix) males displaying at lek, Bergslagen, Sweden, April

Erlend Haarberg / Two Common redpolls (Carduelis flammea) fighting in flight, Norway, March

Erlend Haarberg / Two Arctic terns (Sterna paradisaea) in flight fighting, Iceland, August

Erlend Haarberg / Two Mountain hares (Lepus timidus) fighting, Sor-Trondelag, Norway, April

Erlend Haarberg / Two Mountain hares (Lepus timidus) fighting, Sor-Trondelag, Norway, April

Rolf Nussbaumer / Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...

Ben Hall / Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October

Eric Dragesco / Two Przewalski's / Takhi Horses (Equus ferus prezewalski) fighting, Hustai Nuru National Park, Mongolia

Ernie Janes / Indian rhinoceros (Rhinceros unicornis) charging, captive

Carol Walker / Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009

Carol Walker / Mustang / wild horses, two yearling cremello colts Claro and Cremesso playing with a bay, McCullough Peak herd, Wyoming, USA, June 2008

Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February

Andy Sands / Green Woodpecker (Picus viridis) male foraging on wind-fallen apples, calling in aggression display to deter other birds. Hertfordshire, England, UK, February.

Mark Bowler / Common Squirrel Monkey (Saimiri sciureus ssp. macrodon) in tree, calling, Peru, captive

Gerrit Vyn / Dall sheep (Ovis dalli) rams head butting, one behaviour by which males establish dominance during the Autumn mating season. Denali National Park, Alaska, USA

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder in garden, Berwickshire, Scotland, April

Edwin Giesbers / Two Red fox (Vulpes vulpes) cubs, four months, play fighting, The Netherlands, June

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April

Laurie Campbell / Black-headed gulls (Larus ridibundus) robbing Puffin (Fratercula arctica) of catch, Farne Islands, Northumberland, England, June

Sergey Gorshkov / Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia

Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October. Did you know? Richmond Park deer herds are descended from historic medieval royal hunting stock.

Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, with blocks of flats in the background, Richmond Park, London, England, UK, October

Sergey Gorshkov / African elephant charging (Loxodonta africana) female with young calf, Okavango Delta, Botswana

Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October

Markus Varesvuo / Horned / Slavonian grebe (Podiceps auritus) two individuals fighting in water, Finland May

Tui De Roy / Greater Flamingo (Phoenicopterus ruber) pair in territorial display while two other feed, Galapagos Islands, Ecuador

John Abbott / Eastern Green june beetle (Cotinis nitida) in flight, Texas, USA, October

Kristel Richard / During the Festa do Colete Encarnado (Red Waistcoat Festival), a bull running festival, a traditionally dressed 'cavalheiro' delivers the 'banderilla' mounted on his Lusitano stallion in the bullring...

Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on. Scandinavia, April. 5th, 2010 Euronatur Wildlife photographer competition.

Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on and a Herring Gull (Larus argentus) flies past. Scandinavia, April.

Nick Garbutt / Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November

Nick Garbutt / Adult male Hippopotamuses (Hippopotamus amphibius) posturing in aggressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched togeth...

Sven Zacek / Two Black Grouse (Tetrao tetrix) fighting at a lek. Virumaa, Estonia, Europe, April.

Anup Shah / Caracal (Caracal caracal) two six month kittens play fighting, Masai Mara National Reserve, Kenya, August

Anup Shah / Caracal (Caracal caracal) two six month kittens play fighting, Masai Mara National Reserve, Kenya, August

Andrew Parkinson / 2020VISION / Great crested grebe (Podiceps cristatus) adult aggressively pursues another in a territorial dispute, Derbyshire, UK, March

Terry Whittaker / 2020VISION / Stag beetle (Lucanus cervus) two males displaying aggressive behaviour on oak tree branch at woodland edge, Suffolk, UK, Controlled conditions, July

Roger Powell / Kingfisher (Alcedo atthis) territorial display to warn rivals. Castro Verde, Alentejo, Portugal, April.