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A diver explores Silfra Canyon, a deep fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates at Thingvellir National Park, Iceland. May 2011.  In this phot...
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Alex Mustard / A diver explores Silfra Canyon, a deep fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates at Thingvellir National Park, Iceland. May 2011. In this phot...
Grooved brain coral (Diploria labyrinthiformis) at night with polyps extended to feed, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
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Alex Mustard / Grooved brain coral (Diploria labyrinthiformis) at night with polyps extended to feed, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
Silhouette of Common toad (Bufo bufo) swimming in River Orchy, Highlands, Scotland, UK, April
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Alex Mustard / 2020VISION / Silhouette of Common toad (Bufo bufo) swimming in River Orchy, Highlands, Scotland, UK, April
Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June. 2020VISION Book Plate. Did you know? The largest Basking sha...
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Alex Mustard / 2020VISION / Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June. 2020VISION Book Plate. Did you know? The largest Basking sha...
Grey seal (Halichoerus grypus) swimming amongst kelp, Farne Islands, Northumberland, England, UK, August. 2020VISION Book Plate. Did you know? The 1914 Grey Seal Protection Act ensured this species su...
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Alex Mustard / 2020VISION / Grey seal (Halichoerus grypus) swimming amongst kelp, Farne Islands, Northumberland, England, UK, August. 2020VISION Book Plate. Did you know? The 1914 Grey Seal Protection Act ensured this species su...
Yellowtail tang (Zebrasoma xanthurum) male and female turn belly to belly during spawning to ensure fertilization. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea. This action is very fast...
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Alex Mustard / Yellowtail tang (Zebrasoma xanthurum) male and female turn belly to belly during spawning to ensure fertilization. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea. This action is very fast...
Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies.
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Alex Mustard / Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies.
Yellow tube sponge (Aplysina fistularis) on coral reef, diver exploring in background. Black Rock Drop Off, East End, Grand Cayman, Cayman Islands, British West Indies. Model released.
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Alex Mustard / Yellow tube sponge (Aplysina fistularis) on coral reef, diver exploring in background. Black Rock Drop Off, East End, Grand Cayman, Cayman Islands, British West Indies. Model released.
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt.
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Alex Mustard / Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt.
Green turtle (Chelonia mydas) feeding on Seagrass (Halophila stipulacea), blowing sand out of nostrils, on sea floor. Marsa El Shouna, Marsa Alam, Egypt.
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Alex Mustard / Green turtle (Chelonia mydas) feeding on Seagrass (Halophila stipulacea), blowing sand out of nostrils, on sea floor. Marsa El Shouna, Marsa Alam, Egypt.
Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt.
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Alex Mustard / Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt.
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt.
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Alex Mustard / Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt.
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt.
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Alex Mustard / Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt.
Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean...
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Alex Mustard / Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean...
Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...
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Alex Mustard / Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island. Misool, Raja Ampat, West Papua, Indones...
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Alex Mustard / Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island. Misool, Raja Ampat, West Papua, Indones...
Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean
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Alex Mustard / Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean
California sea lion (Zalophus californianus) pups in an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.
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Alex Mustard / California sea lion (Zalophus californianus) pups in an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.
Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind , in the plankton-rich green waters of La Paz bay, Baja...
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Alex Mustard / Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind , in the plankton-rich green waters of La Paz bay, Baja...
Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
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Alex Mustard / Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
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Alex Mustard / Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
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Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...
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Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...
Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef. This photo shows the larger...
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Alex Mustard / Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef. This photo shows the larger...
Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...
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Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...
Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Veined octopus (Amphioctopus marginatus) shelters inside an old clam shell, with silver-sides in the water above. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Veined octopus (Amphioctopus marginatus) shelters inside an old clam shell, with silver-sides in the water above. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
RF - Silhouette of a California sea lion (Zalophus californianus) swimming through an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific...
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Alex Mustard / RF - Silhouette of a California sea lion (Zalophus californianus) swimming through an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific...
RF - Portrait of Coral hermit crab (Paguritta harmsi) living in a hard coral colony. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Portrait of Coral hermit crab (Paguritta harmsi) living in a hard coral colony. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - A school of Crescent-tail bigeye (Priacanthus hamrur) on a coral reef. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - A school of Crescent-tail bigeye (Priacanthus hamrur) on a coral reef. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
RF - Blue whale (Balaenoptera musculus) diving vertically down into the ocean to feed. Indian Ocean, off Sri Lanka.  (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Blue whale (Balaenoptera musculus) diving vertically down into the ocean to feed. Indian Ocean, off Sri Lanka. (This image may be licensed either as rights managed or royalty free.)
RF - Green sea turtle (Chelonia mydas) Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Green sea turtle (Chelonia mydas) Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image may be licensed either as rights managed or royalty free.)
RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image...
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Alex Mustard / RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image...
Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Portrait of a Giant frogfish (Antennarius commersoni) on a large Yellow elephant ear sponge (Ianthella basta). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Portrait of a Giant frogfish (Antennarius commersoni) on a large Yellow elephant ear sponge (Ianthella basta). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
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Alex Mustard / Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
Portrait of a male short snouted seahorse (Hippocampus hippocampus) in sea oak seaweed (Halidrys siliquosa). Devon, England, United Kingdom. British Isles. English Channel. North East Atlantic Ocean.
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Alex Mustard / Portrait of a male short snouted seahorse (Hippocampus hippocampus) in sea oak seaweed (Halidrys siliquosa). Devon, England, United Kingdom. British Isles. English Channel. North East Atlantic Ocean.
Blue whale (Balaenoptera musculus) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
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Alex Mustard / Blue whale (Balaenoptera musculus) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
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Alex Mustard / Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
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Alex Mustard / Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
Forster's hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt.
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Alex Mustard / Forster's hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt.
Coral (Lobophytum sp.) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata) in background. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
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Alex Mustard / Coral (Lobophytum sp.) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata) in background. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
Hawksbill turtle (Eretmochelys imbricata) unusually feeding on hard coral polyps. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
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Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) unusually feeding on hard coral polyps. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
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Alex Mustard / Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
Carapace of a dead olive Ridley turtle (Lepidochelys olivacea) entangled in discarded fishing gear. Indian Ocean, off Sri Lanka.
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Alex Mustard / Carapace of a dead olive Ridley turtle (Lepidochelys olivacea) entangled in discarded fishing gear. Indian Ocean, off Sri Lanka.
Jellyfish (Catostylus sp.) swims beneath a slick of plastic debris. Indian Ocean off Sri Lanka. Highly commended in the Environmental Report section, Montphoto competition  2019
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Alex Mustard / Jellyfish (Catostylus sp.) swims beneath a slick of plastic debris. Indian Ocean off Sri Lanka. Highly commended in the Environmental Report section, Montphoto competition 2019
Green turtle (Chelonia mydas) descending after breathing at the surface. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
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Alex Mustard / Green turtle (Chelonia mydas) descending after breathing at the surface. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.
Remoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean.
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Alex Mustard / Remoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean.
Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.
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Alex Mustard / Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.
Yellowtail tang (Zebrasoma xanthurum) male courting a female prior to spawning at dusk on a coral reef. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea.
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Alex Mustard / Yellowtail tang (Zebrasoma xanthurum) male courting a female prior to spawning at dusk on a coral reef. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea.
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean.
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Alex Mustard / Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean.
Blue whale (Balaenoptera musculus) diving beneath the surface.  Indian Ocean, Sri Lanka.
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Alex Mustard / Blue whale (Balaenoptera musculus) diving beneath the surface. Indian Ocean, Sri Lanka.
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean.
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Alex Mustard / Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean.
Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean.
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Alex Mustard / Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean.
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean.
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Alex Mustard / Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean.
RF -  Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean. (This image may be licensed either as rights managed or royalty free.)
Hawksbill turtle (Eretmochelys imbricata) pair circling each other above coral reef. Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
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Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) pair circling each other above coral reef. Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
Giant mottled eel (Anguilla marmorata) in river with villagers washing clothes in background. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean.
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Alex Mustard / Giant mottled eel (Anguilla marmorata) in river with villagers washing clothes in background. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean.
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton.  Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
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Alex Mustard / Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Suwanee freshwater turtle (Pseudemys concinna suwanniensis) swimming beneath trees in a freshwater spring. Devil Spring, Ginnie Springs, High Springs, Florida, USA
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Alex Mustard / Suwanee freshwater turtle (Pseudemys concinna suwanniensis) swimming beneath trees in a freshwater spring. Devil Spring, Ginnie Springs, High Springs, Florida, USA
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink. Los I...
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Alex Mustard / California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink. Los I...
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern. Los Islotes, La Paz, Baja California Sur...
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Alex Mustard / Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern. Los Islotes, La Paz, Baja California Sur...
Wave breaking over the rubble zone close to the reef crest of a coral reef. Baa Atoll, Maldives. Indian Ocean.
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Alex Mustard / Wave breaking over the rubble zone close to the reef crest of a coral reef. Baa Atoll, Maldives. Indian Ocean.
American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
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Alex Mustard / American crocodile (Crocodylus acutus) reflected in the surface as it floats, Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea.
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
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Alex Mustard / Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
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Alex Mustard / Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - Spinyhead blenny fish (Acanthemblemaria spinosa) extends out of its hole in a red sponge to snare passing food.  Caribbean Sea (This image may be licensed either as rights managed or royalty free...
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Alex Mustard / RF - Spinyhead blenny fish (Acanthemblemaria spinosa) extends out of its hole in a red sponge to snare passing food. Caribbean Sea (This image may be licensed either as rights managed or royalty free...
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn,  Cayman Islands, Caribbean Sea. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea. (This image may be licensed either as rights managed or royalty free.)
RF - Pod of Atlantic spotted dolphins (Stenella frontalis) swimming over a shallow, sandy seabed. North Bihmini, Bahamas. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF - Pod of Atlantic spotted dolphins (Stenella frontalis) swimming over a shallow, sandy seabed. North Bihmini, Bahamas. (This image may be licensed either as rights managed or royalty free.)
Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell, with plankton swirling around it at night (captured as trails in the long exposure). Bitung, North Sulawesi, Indonesia. Lembeh...
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Alex Mustard / Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell, with plankton swirling around it at night (captured as trails in the long exposure). Bitung, North Sulawesi, Indonesia. Lembeh...
RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. (Th...
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Alex Mustard / RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. (Th...
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
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Alex Mustard / Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
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Alex Mustard / Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
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Alex Mustard / Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
Grey seal (Halichoerus grypus) split level of young seal investigating the camera with its nose and whiskers, close to shore. Lundy Island, Devon, UK, Bristol Channel. August
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Alex Mustard / Grey seal (Halichoerus grypus) split level of young seal investigating the camera with its nose and whiskers, close to shore. Lundy Island, Devon, UK, Bristol Channel. August
Grey seal (Halichoerus grypus) young female opens her mouth and blows bubbles through her nose playfully while she looks up from the seabed. Lundy Island, Devon, UK, Bristol Channel. August. Highly co...
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Alex Mustard / Grey seal (Halichoerus grypus) young female opens her mouth and blows bubbles through her nose playfully while she looks up from the seabed. Lundy Island, Devon, UK, Bristol Channel. August. Highly co...
Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives  after breathing, Lundy Island, Devon, UK, Bristol Channel, August
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Alex Mustard / Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August
Hawksbill turtle (Eretmochelys imbricata) male chomps on soft coral. Jackson Reef, Sinai, Egypt. Strait of Tiran, Red Sea.
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Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) male chomps on soft coral. Jackson Reef, Sinai, Egypt. Strait of Tiran, Red Sea.
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
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Alex Mustard / A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Brandt's cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus) beneath an oil rig. Eureka Rig, Los Angeles, California, USA, September
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Alex Mustard / Brandt's cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus) beneath an oil rig. Eureka Rig, Los Angeles, California, USA, September
Grey seal (Halichoerus grypus) playful young seal above a kelp forest. Lundy Island, Devon, UK Bristol Channel, August
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Alex Mustard / Grey seal (Halichoerus grypus) playful young seal above a kelp forest. Lundy Island, Devon, UK Bristol Channel, August
Grey seal (Halichoerus grypus) split level view of young investigating camera lens, Lundy Island, Devon, UK, Bristol Channel, August
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Alex Mustard / Grey seal (Halichoerus grypus) split level view of young investigating camera lens, Lundy Island, Devon, UK, Bristol Channel, August
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
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Alex Mustard / Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Brandt's cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig. Eureka Rig, Los Angeles, California, United States of Ame...
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Alex Mustard / Brandt's cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig. Eureka Rig, Los Angeles, California, United States of Ame...
California sea lion (Zalophus californianus) adult female swimming through sunbeams in an underwater cave. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.
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Alex Mustard / California sea lion (Zalophus californianus) adult female swimming through sunbeams in an underwater cave. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.
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Alex Mustard / Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
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Alex Mustard / Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea.
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Alex Mustard / Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea.
Harbour seal (Phoca vitulina) in the kelp forest (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
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Alex Mustard / Harbour seal (Phoca vitulina) in the kelp forest (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.