Anup Shah / Chimpanzee {Pan troglodytes} grooming companion Sweetwater Sanctuary, Mahesa, Kenya

Anup Shah / Bonnet macaques {Macaca radiata} grooming one another in tree Periyar NP, Kerala, India

Richard Du Toit / Common Zebra (Equus quagga / burchellii) grooming one another, Gauteng Province, Rietvlei Nature Reserve, South Africa

Bernard Castelein / Khur / Asiatic wild ass (Equus hemionus) two nuzzling each other, Little Rann of Kutch, Gujarat, India

Ann & Steve Toon / Blue duiker (Philantomba / Cephalophus monticola) allogrooming, Mlilwane nature reserve, Swaziland

Pat De La Harpe / Female Impala (Aepyceros melampus) allogrooming in grass, Mala Mala GR, Mpumalanga, South Africa

Bernard Castelein / Impala {Aepyceros melampus} allogrooming, Samburu NP, Kenya

Anup Shah / Mountain gorilla {Gorilla beringei} family group grooming, Parc du Volcans NP, Rwanda

WILLIAM OSBORN / Galah cockatoos allopreening {Eolophus roseicapilla} West Queensland, Australia