Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month hanging upside down in a tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 4...

Danny Green / 2020VISION / Red deer (Cervus elaphus) dominant stag at rut, Bradgate Park, Leicestershire, England, UK, November

Anup Shah / African Lion pride male (Panthera leo) head portrait, low angle, Masai Mara National Reserve, Kenya. February

Suzi Eszterhas / Alpha male Chimpanzee (Pan troglodytes) self grooming. Tropical forest, Western Uganda.

Philip Stephen / Fishing vessel "Ocean Harvest" operating close to the "Beryl Alpha" production platform, 160 miles Northeast of Aberdeen, in the North Sea. March 2010.

Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old, feeding on leaves {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002. Chimps have learnt that certain leaves can be used to tre...

Jim Brandenburg / Arctic Wolf (Canis lupus) alpha male known as Buster, smelling Arctic Poppy (Papaver lapponicum) flowers, Ellesmere Island, Nunavut, Canada

Jim Brandenburg / Timber Wolf (Canis lupus) alpha pair in light snowfall, Minnesota

Jim Brandenburg / Timber Wolf (Canis lupus) alpha female resting as yearling growls at camera, Minnesota

Justine Evans / White handed gibbon, alpha male hanging in tree {Hylobates lar} Khao Yai NP, Thailand 'Cassius'

Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree with dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Alpha male 'Frodo' displays 'keep away' message to begging chimps wanting share of Bushbuck fawn prey {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Ousted alpha male Chimpanzee 'Freud' feeding on mangoes, {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) 'alpha' baby female aged about 1 month sitting in a tree, Tangkoko National Park, Sulawesi, Indonesia.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month sitting playfully with her mother, Tangkoko National Park, Sulawesi, Indonesia.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) 'alpha' male resting against a tree trunk, Tangkoko National Park, Sulawesi, Indonesia.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month sitting in her mother's arms, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month sitting with her mother in a tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged 1-2 months playing with older juveniles, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month playing with a coconut shell, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' male sitting in the fork of a tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month attempting to feed on fruits in a tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month sitting on log, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month playing in a tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Western chimpanzee (Pan troglodytes verus) female 'Jire' aged 52 years being groomed by alpha male 'Foaf' aged 30 years, whilst she feeds on palm oil fruit, Bossou Forest, Mont Nimba, Guinea. Januar...

Anup Shah / Western chimpanzee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years looking curiously at female 'Jire' aged 52 years as she feeds on palm oil fruits, Bossou Forest, Mont Nimba, Guinea. Januar...

Anup Shah / Western chimpanzee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years grooming young male 'Peley' aged 12 years, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) female 'Fanle' aged 13 years with suckling male infant 'Flanle' aged 3 years being groomed by alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, G...

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years aggressive towards alpha male 'Foaf' aged 30 years because he is interested in his mother 'Fanle' aged 13 years, Bossou F...

Anup Shah / Western chimpanzee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years walking into a clearing with young male 'Jeje' aged 13 years sitting behind, Bossou Forest, Mont Nimba, Guinea. January 201...

Anup Shah / Western chimpanzee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years making a 'Pant Hoot' volcalisation to make contact with other community members, Bossou Forest, Mont Nimba, Guinea. Decembe...

Anup Shah / Western chimpanzee (Pan troglodytes verus) female 'Fanle' aged 13 years with her infant son 'Flanle' aged 3 years on her back sitting with alpha male 'Foaf' aged 30 years who is cracking open palm o...

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years who then 'kisses' him on the forehead to diffuse the situatio...

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 3...

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 2...

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 1...

Anup Shah / Western chimpanzee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years grooming female 'Jire' aged 52 years while she feeds on palm oil fruits, Bossou Forest, Mont Nimba, Guinea. January 2011.

Fiona Rogers / Western chimpazee (Pan troglodytes verus) female 'Jire' aged 52 years being groomed by alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011.

Fiona Rogers / Western chimpazee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years looking curiously at female 'Jire' aged 52 years as she feeds on palm oil fruits. Bossou Forest, Mont Nimba, Guinea. January 2...

Fiona Rogers / Western chimpazee (Pan troglodytes verus) female 'Fanle' aged 13 years with suckling male infant 'Flanle' aged 3 years being groomed by alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guin...

Fiona Rogers / Western chimpazee (Pan troglodytes verus) female 'Fanle' aged 13 years with suckling male infant 'Flanle' aged 3 years being groomed by alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guin...

Fiona Rogers / Western chimpazee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years charging, Bossou Forest, Mont Nimba, Guinea. December 2010.

Fiona Rogers / Western chimpazee (Pan troglodytes verus) alpha male 'Foaf' aged 30 years vocalising whilst sitting, Bossou Forest, Mont Nimba, Guinea. January 2011.

Danny Green / European Wolf (Canis lupus) alpha male walking beside water. Finland, July.

Danny Green / European Wolf (Canis lupus) alpha male in meadow. Finland, July.

Danny Green / European Wolf (Canis lupus) alpha male by water, Finland, July.

Danny Green / European Wolf (Canis lupus) alpha female on grassland. Finland, July.

Danny Green / European Wolf (Canis lupus), alpha female walking in grassland. Finland, July.

Tony Phelps / Chacma baboon (Papio hamadryas ursinus), alpha male drinking from vlei. deHoop, Western Cape, South Africa, September.

Tony Phelps / Chacma Baboon (Papio hamadryas ursinus), alpha male walking along water's edge. deHoop Vlei, Western Cape, South Africa, September.

Jabruson / Agile Mangabey (Cercocebus agilis) alpha male profile. Bai Hokou, Dzanga-Ndoki National Park, Central African Republic.

Jabruson / Western lowland gorilla (Gorilla gorilla gorilla) silverback 'Makumba' portrait, leader of tourist habituated gorilla family group, Bai Hokou, Dzanga-Ndoki National Park, Central African Republic.

Jabruson / Agile mangabey (Cercocebus agilis) female resting with alpha male close-by. Bai Hokou, Dzanga-Ndoki National Park, Central African Republic.

Jabruson / Agile Mangabey (Cercocebus agilis) alpha male resting with female close-by. Bai Hokou, Dzanga-Ndoki National Park, Central African Republic.

Jabruson / Agile mangabey (Cercocebus agilis) alpha male resting. Bai Hokou, Dzanga-Ndoki National Park, Central African Republic.

Christophe Courteau / African wild dog in submissive pose to alpha pregnant female Okavango

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Male Chimpanzees fighting (Pan troglodytes) heavily injured alpha male 'Pim', Mahale Mountains National Park, Tanzania, East Africa

Philip Stephen / The Cormorant Alpha production platform, situated in the North Sea 70 miles north east of the Shetland Islands, May 2007

Philip Stephen / Offloading containers at the Beryl alpha platform, North Sea, October 2009.

Rick Tomlinson / "Alpha Romeo" during Coastal Race 1, Rolex Maxi Cup, Porto Cervo, Sardinia, Italy. 7 September 2009.

Konrad Wothe / Chimpanzee (Pan troglodytes) males after deadly fight sitting around dead alpha male, Mahale Mountains National Park, Tanzania

Konrad Wothe / Chimpanzee (Pan troglodytes) males killing alpha male, Mahale Mountains National Park, Tanzania

Ann & Steve Toon / African wild dog (Lycaon pictus) alpha female rolling in dirt, Kruger National Park, South Africa, May.

Wild Wonders of Europe / Lesniewski / Red Deer (Cervus elaphus) females, a harem belonging to an alpha male. Bieszczady, Carpathian Mountains, Poland, September.

Florian Möllers / Alpha-female Chimpanzee (Pan troglodytes schweinfurthii) "Nambi" grooming her daughter "Night" (3 years) in tree. Budongo Forest Reserve, Masindi, Uganda, Africa. December

Edwin Giesbers / Orang utan (Pongo pygmaeus) head portrait of dominant male called Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered

Edwin Giesbers / Orang utan (Pongo pygmaeus) head portrait of dominant male Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered

Michael Quinton / Gray Wolf (Canis lupus) alpha wearing tracking collar leading pack, Denali National Park, Alaska

Pete Oxford / Southern Elephant Seal (Mirounga leonina) Bull vocalising after fighting for control of the harem. Sea Lion Island. South of mainland east Falkland Island. Falkland Islands

Suzi Eszterhas / Alpha male Chimpanzee (Pan troglodytes) reclining on forest floor. Tropical forest, Western Uganda.

Justine Evans / Alpha female Bornean Muller's / Grey gibbon (Hylobates muelleri) holding fruit in foot hanging from a fruiting fig tree, Danum Valley, Sabah, Borneo, Endangered species

Justine Evans / Alpha female Grey gibbon (Hylobates muelleri) leaping from a fruiting strangler-fig tree, Danum Valley, Sabah, Borneo, Endangered species

Justine Evans / Alpha female Bornean Muller's / Grey gibbon (Hylobates muelleri) feeding on fruit in a strangler fig tree, while hanging from one arm, Danum Valley, Sabah, Borneo, Endangered species

Justine Evans / Alpha-male White-handed gibbon (Hylobates lar) sitting on branch observing a neighboring gibbon family on the border of his territory, "Chai" a member of study group "C", Khao Yai National Park, Thail...

Justine Evans / Alpha male White-handed gibbon (Hylobates lar) hanging from strangler fig, "Chai" a member of study group "C", Khao Yai National Park, Thailand, Endangered species

Justine Evans / Alpha male White-handed gibbon (Hylobates lar) in tree observing a neighboring gibbon family on the border of his territory, "Chai" a member of study group "C", Khao Yai National Park, Thailand, Endan...

Justine Evans / Alpha female White-handed gibbon (Hylobates lar) sitting in tree, "Cassandra" a member of study group "C", Khao Yai National Park, Thailand, Endangered species

Suzi Eszterhas / Chimpanzee (Pan troglodytes) alpha male, western Uganda

Suzi Eszterhas / Chimpanzee (Pan troglodytes) alpha male being groomed by brother, western Uganda

Suzi Eszterhas / Chimpanzee (Pan troglodytes) alpha male and brother showing hand clasp grooming behavior, western Uganda

Suzi Eszterhas / Chimpanzee (Pan troglodytes) alpha male, western Uganda

Suzi Eszterhas / Chimpanzee (Pan troglodytes) alpha male, Ngamba Island Chimpanzee Sanctuary, Uganda

Angelo Gandolfi / European Grey wolf (Canis lupus italicus) in snow, captive, Alpha Park, Parc National du Mercantour, Alps, Southern France

Angelo Gandolfi / European Grey wolf (Canis lupus italicus) walking through snow, captive, Alpha Park, Parc National du Mercantour, Alps, Southern France

Angelo Gandolfi / Visitor in the wolf watching shed at Alpha Park, Parc National du Mercantour, Alps, Southern France

Angelo Gandolfi / European Grey wolf (Canis lupus italicus) clawing at tree with dead robin in its mouth, captive, Alpha Park, Parc National du Mercantour, Alps, Southern France

Angelo Gandolfi / European Grey wolf (Canis lupus italicus) walking through snow, captive, Alpha Park, Parc National du Mercantour, Alps, Southern France