Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Sergio Hanquet / Arrow crab (Sternorhynchus lanceolatus). Tenerife, Canary Islands.

Guy Edwardes / Marbled white (Melanagria galathea), two butterflies on Pyramidal orchid (Anacamptis pyramidalis). Badbury Rings, Wimborne Minster, Dorset, England, UK. June.

Heather Angel / Honey bee (Apis mellifera) forages on pollen in Apple (Malus domestica) flower, collecting pollen in pollen basket. In garden, Surrey, England, UK. May.

Alex Mustard / Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...

Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Fabio Pupin / Red swamp crayfish (Procambarus clarcki), invasive species, captive, Italy, July.

Nigel Cattlin / Insect / invertebrate house / bug hotel with various refuge options for hiding / over-wintering, WWT, West Sussex, UK, July.

John Chan / Crenulated fiddler crab (Uca crenulata) waving its major claw at another crab, part of a courtship display, Southern California, USA, June.

John Chan / Crenulated fiddler crab (Uca crenulata) feeding on detritus, Southern California, USA, June.

John Shaw / Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May.

Shane Gross / Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp). Anilao, Philippines.

Andy Rouse / Sally lightfoot crab, (Grapsus grapsus), feeding on marine iguana parasites and dead skin, Galapagos

Georgette Douwma / Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia.

Nick Hawkins / European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July.

SCOTLAND: The Big Picture / Close up of a Long clawed squat lobster (Munida rugosa) in Loch Carron, January 2016.

Steven David Miller / Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.

Juergen Freund / Wide-angle timelapse of Sand bubbler crabs (Dotillidae) feeding, filtering detritus and plankton from sand and discarding the processed sand as pellets, which cover the beach, Wet Tropics World Herita...

Juergen Freund / Sand bubbler crabs (Dotillidae) feeding, filtering detritus and plankton from sand and discarding the processed sand as pellets, which cover the beach, Wet Tropics World Heritage Area, Cape Tribulatio...

Juergen Freund / Sand bubbler crab (Dotillidae) feeding, filtering detritus and plankton from sand and discarding the processed sand as pellets, which cover the beach, Wet Tropics World Heritage Area, Cape Tribulation...

Remi Masson / European crayfish (Astacus astacus) in a lake. Alps, Switzerland, July.

David Fleetham / Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp), Philippines.

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or defenc...

Nick Upton / Common prawns (Palaemon serratus) and a Rock goby (Gobius paganellus) feeding on a dead Montagu's crab (Xantho hydrophilus) in a rockpool, Cornwall, England, UK, September.

Nick Upton / Rock goby (Gobius paganellus) stealing food from a scavenging Common prawn (Palaemon serratus) in a rockpool, Cornwall, England, UK, September.

Sue Daly / Common lobster (Homarus gammarus) shovelling gravel out of a hole, Sark, British Channel Islands, UK, August.

Sue Daly / Long legged spider crab (Macropodia) walking across the seabed, Sark, British Channel Islands, UK, June.

Ammonite / Close up of a Common snail (Cornu aspersum) moving, with Woodlice (Oniscidea). Controlled conditions.

David Shale / Pram bug (Phronima) swimming, Mid-Atlantic Ridge. Deep sea species.

Ingo Arndt / Rear view of Horseshoe crab (Limulus polyphemus) with trail in sand, Delaware Bay, Delaware, USA, May

Ingo Arndt / Christmas Island Red Crab (Gecarcoidea natalis) blurred view of vast number of individuals moving along their annual migration route, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Robber / Coconut Crab (Birgus latro) close up of head and legs, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) crossing closed road, during annual migration, Christmas Island, Indian Ocean, Australian Territory, November 2009.

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) crossing road in huge numbers during annual migration, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) arriving in huge numbers, at coast for spawning, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) gatthered together on beach the day before spawning, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) massing on coastal rocks, the morning after spawning, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Christmas Island Red Crabs (Gecarcoidea natalis) shortly before sunrise sitting on rock ready for spawning, Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Four spotted libellula dragonfly / (Libellula quadrimaculata) adult freshly emerged from larval case, Germany.

Ingo Arndt / Honey Bee (Apis mellifera) adult hatching out of brood cell, Germany.

Ingo Arndt / Honey Bee (Apis mellifera) opening brood cell before hatching, Germany. Winner of Fritz Polking portfolio prize, GDT 2012 competition.

Ingo Arndt / Bluebottle Fly (Calliphora erythrocephala) hatching from coccoon, Germany.

Ingo Arndt / Monarch butterflies (Danaus plexippus) gathered to drink, overwintering colony, Michoacan, Mexico

Ingo Arndt / China mark moth (Nymphula nympheata) on water surface, having just hatched underwater

Ingo Arndt / Close-up of eyes on wing of Common Morpho butterfly (Morpho peleides) Costa Rica

Ingo Arndt / Cloud of Monarch butterflies (Danaus plexippus) flying, overwintering colony, Michoacan, Mexico

Ingo Arndt / Monarch butterflies (Danaus plexippus) flying amongst trees, overwintering colony, Michoacan, Mexico

Ingo Arndt / Painted lady butterfly (Vanessa cardui) in flight, Germany

Ingo Arndt / Close-up of Owl butterfly (Caligo memnon) face, Central America (Captive)

Ingo Arndt / Pupa of Common sargeant butterfly (Athyma perius)

Ingo Arndt / Monarch butterfly (Danaus plexippus) hanging from a blade of grass, silhouetted against the full moon, East coast USA.

Ingo Arndt / Monarch butterflies (Danaus plexippus) flying in warmth of midday sun, overwintering colony in Michoacan, Mexico

Ingo Arndt / Monarch butterflies (Danaus plexippus) drinking from pool of water, overwintering colony, Michoacan, Mexico

Ingo Arndt / Male Monarch butterfly (Danaus plexippus) feeding on nectar of Goldenrod flowers, East coast, USA

Ingo Arndt / Black garden ants {Lasius niger} feeding on cricket Germany, Europe

Tony Wu / Humpback whale (Megaptera novaeangliae) adult female at rest. The tubercles on the whale's head are clearly visible, as are the barnacles (Coronula diadema), whale lice (Cyamus boopis), and hairs on t...

Doc White / California sea otter (Enhydra lutris) feeding on Northern kelp crab, Monterey Bay, California, USA, Eastern Pacific Ocean, May

Lorraine Bennery / Knapweed fritillary butterflies (Melitaea phoebe) on Green winged orchid (Anacamptis morio), Grands Causses Regional Natural Park, France, May.

Lorraine Bennery / Geometrid moth caterpillar (Geometridae sp.) moving along a twig in classic pose, Corbieres, France, May.

Clay Bolt / Central bumble bee (Bombus centralis) pollinating wildflowers in the Bridger Range, Bozeman, Montana. August.

John Abbott / Paper wasp (Polistes metricus) flying from flower, Tuscaloosa County, Alabama, USA Controlled conditions. September

James Dunbar / Leaf cutter ants (Atta sp.) communicating. Guadeloupe National Park, Guadeloupe, Leeward Islands.

Nick Hawkins / Leaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica.

Alex Hyde / Fishing spider (Trechalea) waiting to catch a Killifish (Cyprinodontiformes) in a rainforest stream at night, Central Caribbean foothills, Costa Rica.

Alex Hyde / Leaf-cutter ant (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica

Alex Hyde / Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica. Photographed in mobile field studi...

Alex Hyde / Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica. Photographed in mobile field studi...

Alex Hyde / Jumping spider (Salticidae) hunting among vegetation, San Jose, Costa Rica.

Andy Sands / Jumping spider (Evarcha arcuata) on edge of evening primrose flower, Surrey, England, UK, August

Stephen Dalton / Horse fly (Tabanus sp) male taking off from poppy seed-head (Papaver rhoeas) Sussex, UK

Floris van Breugel / Monarch butterflies (Danaus plexippus) over winter on Eucalyptus and Monterey Cypress trees near Pismo Beach, San Luis, California, USA December.. When temperatures warm up enough, they will embark on...

Pal Hermansen / Garden cross spider (Araneus diadematus) on web, Norway.

Pal Hermansen / Hoverfly in flight, backlight, Norway, September.

Bence Mate / Leaf cutter ants (Atta sp) carrying piece of leaf, Costa Rica.

Bence Mate / Leaf cutter ants (Atta sp) carrying pieces of fern, reflected in water, Costa Rica.

Bence Mate / Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica.

Bence Mate / Leafcutter ants (Atta sp) colony harvesting pieces of a banana leaf, Costa Rica.

Bence Mate / Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Leaf cutter ants (Atta sp) silhouetted carrying plant matter, Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica.

Bence Mate / Leaf cutter ant (Atta sp) carrying pink flower segment, Santa Rita, Costa Rica.

Bence Mate / Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Costa Rica.

Jan Hamrsky / Brine shrimp or monkey shrimp (Artemia salina) female. Controlled conditions.

Jan Hamrsky / Diving beetle larva (Hyphydrus ovatus), Europe, June. Controlled conditions.

Jan Hamrsky / Diving beetle (Haliplus ruficollis), Europe, June. Controlled conditions.

Jan Hamrsky / Diving beetle (Colymbetes fuscus) close up head portrait. Europe, June. Controlled conditions.

Jan Hamrsky / Summer mayfly nymph (Siphlonurus lacustris), Europe, May. Controlled conditions.

Jan Hamrsky / Summer mayfly nymphs (Siphlonurus lacustris), Europe, May. Controlled conditions.

Jan Hamrsky / Case-building caddisfly larva (Trichoptera), Europe, June. Controlled conditions.

Jan Hamrsky / Hacklegill mayfy nymph (Potamanthus luteus), Europe, July. Controlled conditions.

Alex Hyde / Nine-spotted moth (Amata phegea) feeding from a Mountain arnica flower (Arnica montana) Triglav National Park, Julian Alps, Slovenia. July.

Alex Hyde / Harvestman (Leiobunum sp.) Triglav National Park, Julian Alps, Slovenia, July.

Alex Hyde / European House Centipede (Scutigera coleoptrata) found on the ceiling of a limestone cave. Photographed on a white background in mobile field studio. Plitvice Lakes National Park, Croatia. November.

Alex Hyde / Close up of head of European House Centipede (Scutigera coleoptrata) in limestone cave. Plitvice Lakes National Park, Croatia. November.

Alex Hyde / Painted lady butterfly (Vanessa cardui) on flower, Peak District National Park, Derbyshire, UK. September.