Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

John Shaw / Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with with creche of huddling chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Nick Upton / White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...

Nick Upton / White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.

Nick Upton / Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...

Nick Upton / Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...

Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, males huddling on sea ice during polar night, incubating eggs, ice shelf in background. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled during polar night, males incubating eggs. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) males in breeding colony huddling during winter storm. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddling, males incubating eggs, on sea ice under full moon in polar night. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony on sea ice, ice shelf in background. Atka Bay, Antarctica. August.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) males in breeding colony incubating eggs, huddling during winter storm. Penguins on outside repositioning themselves to gain protection from weather. Atka Bay, A...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, mainly males brooding eggs, huddled together on sea ice. Atka Bay, Antarctica. June.

Stefan Christmann / Cameraman Lindsay McCrae filming Emperor penguin (Aptenodytes forsteri) colony for BBC Dynasties Penguin programme. Chicks are on sea ice whilst parents fish at sea. Atka Bay, Antarctica. December 201...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding over ice, returning to form breeding colony. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding over sea ice, returning to form breeding colony. Iceberg in background. Atka Bay, Antarctica. April 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) walking across crack in sea ice during return to breeding colony. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding across sea ice to form breeding colony. Atka Bay, Antarctica. April 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) sliding across sea ice in journey to breeding colony. Sastrugi in ice formed by wind erosion. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) forming breeding colony at start of winter. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, adults brooding chicks. Atka Bay, Antarctica. September 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), breeding colony with individuals in foreground walking to / from open water. Atka Bay, Antarctica. September 2017.

Paul Souders/ Worldfoto / Lesser Flamingo (Phoenicopterus minor) flock nesting on island, Lake Natron, Tanzania

Paul Souders/ Worldfoto / Lesser Flamingo (Phoenicopterus minor) nesting colony, Lake Natron, Tanzania

Philippe Clement / Wide-angle shot of Northern gannets (Morus bassanus) nesting in a breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) in breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) sitting on nests made with plastic, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) preening, sitting on nests made with plastic, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) preening at breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Great cormorant (Phalacrocorax carbo) bringing twig to nest in breeding colony, Belgium, March.

Philippe Clement / Common guillemots (Uria aalge) and Razorbills (Alca torda) swimming and fishing at sea, Fowlsheugh RSPB Reserve, Aberdeenshire, Scotland, UK, May.

Philippe Clement / Razorbill (Alca torda) looking around on sea cliff top at seabird colony in spring, Fowlsheugh RSPB Reserve, Aberdeenshire, Scotland, UK, May

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Finalist in the Portfolio Category of the Terre Sauvage Nature Images Awards...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick under the wing of protective parent, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June.Finalist in the Portfolio Category...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick age three days, standing in nest, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category o...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick creche in front of attentive adult flamingo group, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Por...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick age 5 days, walking around nest, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) sleeping in the breeding colony, brooding egg, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of the Terr...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) tending to newborn chick while another feed two day old chick, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in t...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) chick group walking around the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of the Ter...

Claudio Contreras / Aerial view of Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May, Finalist in the Portfolio Category of the Terre Sauvage Nature...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) preening chick in the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June Finalist in the Portfolio Category of the Terre Sauvage...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) building up mud whilst brooding egg, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) feeding three day old chick at breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, June, Finalist in the Portfolio Category of t...

Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) rolling egg to side in the nest, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, June, Finalist in the Portfolio Category of the Terre Sauvag...

Tui De Roy / Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos

Frederique Olivier / Close-up of a group of Emperor penguin (Aptenodytes forsteri) chicks huddled together in a blizzard, covered in snow, Adelie Land, Antarctica, January.

Frederique Olivier / Emperor penguin (Aptenodytes forsteri) chicks huddled together during a blizzard, Adelie Land, Antarctica, January.

Frederique Olivier / Group of Adelie penguins (Pygoscelis adeliae) destroying an egg with a chick inside, Adelie Land, Antarctica, January.

Frederique Olivier / Close-up of a Black browed albatross (Thalassarche melanophris) preening its chick, chick looks up panting, thermoregulating, West Point Island, Falkland Islands.

Frederique Olivier / Southern rockhopper penguin (Eudyptes chrysocome) at nest in breeding colony, stretching and knocking chick over, New Island, Falkland Islands.

Frederique Olivier / Southern rockhopper penguin (Eudyptes chrysocome) preening chick then itself, New Island, Falkland Islands.

Frederique Olivier / Black browed albatross (Thalassarche melanophris) chick in nest, interacting with a Southern rockhopper penguin (Eudyptes chrysocome) chick, New Island, Falkland Islands.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) moving a feather in its nest, Aitcho Island, South Shetland Islands.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) adding a stick to its nest and fighting with a neighbouring pair, Aitcho Island, South Shetland Islands.

Frederique Olivier / Panning shot of a Chinstrap penguin (Pygoscelis antarcticus) breeding colony, Aitcho Island, South Shetland Islands.

Peter Bassett / Pair of Gentoo penguins (Pygoscelis papua) courting and attempting to mate, Neko Harbour, Andvord Bay, Graham Land, Antarctica.

Peter Bassett / Panning shot of a King penguin (Aptenodytes patagonicus) breeding colony, Salisbury Plain, South Georgia.

Peter Bassett / King penguin (Aptenodytes patagonicus), Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) preening in breeding colony, with mountains in the background, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) entering and leaving the sea near their breeding colony, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) in a breeding colony, with mountains in the background, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) walking towards and entering the sea near their breeding colony, Salisbury Plain, South Georgia.

Frederique Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Tui De Roy / Snares Crested penguin (Eudyptes robustus) group walking up granite slope into Oleander forest, Station Point colony. Snares Island, New Zealand Sub-Antarctic Islands.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) colony, with icebergs and pack ice in the background, Antarctica.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) displaying in the middle of a colony, Antarctica.

Claudio Contreras / Great Frigatebird (Fregata minor) departing from nest, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Isl...

Jack Dykinga / Wrecked ship with cormorant colony, on Namibia's skeleton coast, Dorob National Park, Namibia. July 2013.

Neil Aldridge / African penguins (Spheniscus demersus) preening themselves on rocks at the Boulders Beach colony near Cape Town, South Africa.

David Pattyn / Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Republic of Ireland, June

Hiroya Minakuchi / King Penguin (Aptenodytes patagonicus) colony, South Georgia Island

Mark MacEwen / Snares island crested penguin (Eudyptes robustus) colony in forest, Snares Island, New Zealand.

Mark MacEwen / Snare's island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand.

Dave Watts / Hermit ibis (Geronticus eremita) colony, near Tamri, Morocco. Critically endangered.

Angelo Gandolfi / White stork (Ciconia ciconia), nesting colony to provide alternative nesting area whilst the building they normally nest on is under repair. Extremadura, Spain, April.

David Tipling / Northern gannet (Sula bassana) colony, Hermaness, Unst, Shetland Scotland, UK, April.

Espen Bergersen / Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) colony at sunrise. Grytviken, South Georgia Island.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) colony with storm approaching. Grytviken, South Georgia Island.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) and Southern elephant seal (Mirounga leonina) colony at sunrise. Grytviken, South Georgia Island.

Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) colony huddling for warmth, Queen Maud Land, Antarctica

Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) colony huddling for warmth, Queen Maud Land, Antarctica

Mark Jones / Snares-crested penguin (Eudyptes robustus) group climbing steep route to colony. Snares Island, New Zealand Subantarctic Islands. Endemic. Vulnerable species.

Bryan and Cherry Alexander / Emperor penguins (Aptenodytes forsteri) colony, with Snow petrel (Pagodroma nivea) flying past and dark sky. Atka Bay. Antarctica

Cheryl-Samantha Owen / Cape gannet (Morus capensis) colony, Bird Island Nature Reserve, Lambert's Bay, West Coast, South Africa.