Ann & Steve Toon / Ostrich (Struthio camelus) silhouetted in dust storm. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa. Commended in the Bird Photographer of the Year...

Andres M. Dominguez / Hawfinch (Coccothraustes coccothraustes) perched on branch amongst Hawthorn (Crataegus sp) berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February.

Claudio Contreras / California condor (Gymnogyps californianus), portrait. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico.

Claudio Contreras / California condor (Gymnogyps californianus), two peched in tree, adult and juvenile. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico.

Jack Dykinga / Greater roadrunner (Geococcyx californianus) running to feed brood with Lizard prey in beak. Catalina State Park, Sonoran Desert, Arizona, USA. June.

George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.

George Sanker / Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.

Staffan Widstrand / African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Staffan Widstrand / Tawny eagle (Aquila rapax) in flight, about to land, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Oscar Dewhurst / Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September

Lucas Bustamante / Anhinga (Anhinga anhinga). Yasuni National Park, Orellana, Ecuador

Lucas Bustamante / Zigzag Heron (Zebrilus undulatus) camouflaged in branches while nesting. Yasuni National Park, Ecuador

Lucas Bustamante / Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) juveniles and a few adults kept temporarily in an enclosure for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) juveniles and a few adults kept temporarily in an enclosure for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Volunteers rounding up Greater flamingo (Phoenicopterus ruber) juveniles for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) flock silhouetted in wetland, Fuente de Piedra lagoon, Malaga, Spain. August.

Bernard Castelein / Yellow-rumped cacique (Cacicus cela) calling, Madidi NP, Bolivia

Bernard Castelein / Indian Flying Fox (Pteropus giganticus) male roosting in tree, Keoladeo NP, Bharatpur, India

Bernard Castelein / Rhesus macaque (Macaca mulatta), juveniles playing in tree, Keoladeo NP, Bharatpur, India

Bernard Castelein / Spotted owlet ( Athena brama), Keoladeo NP, Bharatpur, India

Bernard Castelein / Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India

Bernard Castelein / Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India

Bernard Castelein / Intermediate egret (Egrettta intermedia), Keoladeo NP, Bharatpur, India

Steven David Miller / Rainbow lorikeet (Trichoglossus moluccanus) on a branch, Cania Gorge National Park, Queensland, Australia. September.

Steven David Miller / Olive-backed sunbird (Nectarinia jugularis) female returning to her nest with nesting material, Cape Hillsborough National Park, Queensland, Australia, September.

Steven David Miller / Olive-backed Sunbird (Nectarinia jugularis) female in her nest, Cape Hillsborough National Park, Queensland, Australia, September.

Marie Read / Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her and crossing his bill with hers...

Marie Read / Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January.

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

David Pattyn / Great crested grebe (Podiceps cristatus) adult with freshly caught fish. Tench (Tinca tinca) to feed to its young chick, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June

David Pattyn / Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July

David Pattyn / Great crested grebe (Podiceps cristatus) adult with young on its back and in the water, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June

David Pattyn / Great crested grebe (Podiceps cristatus) adult with young on its back, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June

David Pattyn / Great crested grebe (Podiceps cristatus) adults courting and greeting each other to confirm their bond. Even with the young chicks already out they often show courtship behavior to each other, Valkenh...

David Pattyn / Mute swan (cygnus olor) family, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June

David Pattyn / Golden eagle (Aquila chrysaetos) in flight in front of a vertical rocky mountain side, Gran Paradiso National Park, Aosta Valley, Italy, May

David Pattyn / Plate-billed Mountain-Toucan (Andigena laminirostris), Bellavista private reserve, Mindo cloud forest, Pichincha, Ecuador, July

David Gallan / Slow motion clip of a Wedge-tailed eagle (Aquila audax) landing in nest, Mimosa National Park, New South Wales, Australia , September.

George Sanker / Greater yellowlegs (Tringa melanoleuca) standing on rock. Acadia National Park, Maine, USA. September.

John Waters / Large murmuration of Common starlings (Sturnus vulgaris) swirling over reedbed roost at sunset, Ham Wall RSPB Reserve, Somerset, England, UK, January.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.

Jed Weingarten / Wild Wonders of China / Mrs Gould's sunbird (Aethopyga gouldiae) perched on branch, Tangjiahe Nature Reserve, Sichuan Province, China.

Jed Weingarten / Wild Wonders of China / Temminck's tragopan (Tragopan temminckii) Tangjiahe Nature Reserve, Sichuan province, China.

Pedro Narra / Cuban emerald hummingbird (Chlorostilbon ricordii) Guanahacabibes Peninsula National Park, Cuba

Pedro Narra / Bee hummingbird (Mellisuga helenae) hovering in front of flower, Guanahacabibes Peninsula National Park, Cuba

Pedro Narra / Bee hummingbird (Mellisuga helenae) feeding from flower, Guanahacabibes Peninsula National Park, Cuba

SCOTLAND: The Big Picture / Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK.

Ann & Steve Toon / Spotted eagle owl (Bubo africanus) subadult at nigh by water, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Kerstin Hinze / Red crossbill (Loxia curvirostra) feeding from a spruce cone, Bavarian Forest National Park, Germany, October.

Kerstin Hinze / Male Capercaillie (Tetrao urogallus) displaying, Berchtesgaden National Park, Bavarian Alps, Germany, April.

Kerstin Hinze / Male Capercaillie (Tetrao urogallus) displaying, Berchtesgaden National Park, Bavarian Alps, Germany, April.

Kerstin Hinze / Dipper (Cinclus cinclus) foraging in a stream, Harz National Park, Saxony-Anhalt, Germany, March.

Kerstin Hinze / Male Black woodpecker (Dryocopus martius) drilling a hole in a tree, hunting for insects, Bavarian Forest National Park, Bavaria, Germany, February.

Philippe Clement / Wide-angle shot of Northern gannets (Morus bassanus) nesting in a breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) in breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) sitting on nests made with plastic, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) preening, sitting on nests made with plastic, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

Philippe Clement / Northern gannets (Morus bassanus) preening at breeding colony, Hermaness NNR, Unst, Shetland Islands, Scotland, UK, May.

John Chan / Osprey (Pandion haliaetus) feeding on a fish, perched on a dead branch, Bolsa Chica Ecological Reserve, California, USA, October.

Richard Du Toit / Red-billed Oxpecker (Buphagus erythrorhynchus) on Burchell's Zebra (Equus burchellii), Mkhuze Game Reserve, KwaZulu-Natal, South Africa

Richard Du Toit / Northern Giraffe (Giraffa camelopardalis) with Red-billed Oxpeckers (Buphagus erythrorhynchus) feeding on parasites in symbiotic relationship, Mkhuze Game Reserve, KwaZulu-Natal, South Africa

Richard Du Toit / Purple-crested Turaco (Tauraco porphyreolophus), Itala Game Reserve, KwaZulu-Natal, South Africa

David Tipling / Eurasian magpies (Pica pica) five gathering prior to going to roost Hortobagy National Park, Hungary January

Nick Garbutt / Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru.

Nick Garbutt / Red-and-green macaw (Ara chloropterus) flock flying in front of clay lick. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border.

Staffan Widstrand / Wild Wonders of China / Golden pheasant (Chrysolophus pictus) male jumping, Yangxian Biosphere Reserve, Shaanxi, China

Staffan Widstrand / Wild Wonders of China / Golden pheasant (Chrysolophus pictus) male displaying in grass in Yangxian Nature Reserve, Shaanxi, China, September.

Staffan Widstrand / Wild Wonders of China / Crested ibis (Nipponia nippon) flying, Yangxian Nature Reserve, Shaanxi, China, September.

Staffan Widstrand / Wild Wonders of China / Red jungle fowl (Gallus gallus) Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April.

Bence Mate / European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February.

Chris Gomersall / Large flock of Red knot (Calidris canutus) moving at high tide roost, in winter plumage, Snettisham RSPB Reserve, Norfolk, England, UK, September.

Guy Edwardes / Saddle-billed stork (Ephippiorhynchus senegalensis), Moremi Game Reserve, Botswana.

Andy Trowbridge / Kea (Nestor notabilis) juvenile in flight. Arthur's Pass National Park, South Island, New Zealand. Endangered Species.

Juergen & Christine Sohns / Black-footed Penguin (Spheniscus demersus) pair on beach, Betty's Bay, Stony Point Nature Reserve, South Africa

Konrad Wothe / Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June.

George Sanker / Osprey (Pandion haliaetus) with two Alewife (Alosa pseudoharengus) just caught in the Atlantic Ocean. Acadia National Park, Maine, USA. June.

Jaymi Heimbuch / Yellow-billed Oxpecker (Buphagus africanus) picking parasites off of Cape Buffalo (Syncerus caffer), Masai Mara, Kenya

Oscar Dewhurst / Long-crested eagle (Lophaetus occipitalis) perched on bush. Selenkay Conservancy, Amboseli National Park, Kenya.

Oscar Dewhurst / Nubian woodpecker (Campethera nubica) foraging on tree trunk. Ol Kinyei Conservancy, Masai Mara National Reserve, Kenya.

Oscar Dewhurst / Long-crested eagle (Lophaetus occipitalis) perched on tree. Selenkay Conservancy, Amboseli National Park, Kenya.

Oscar Dewhurst / Mountain bluebird (Sialia currucoides). Yellowstone National Park, Wyoming, USA. September.

Oscar Dewhurst / Trumpeter swan (Cygnus buccinator) pair on bank. Grand Teton National Park, Wyoming, USA. September.

Oscar Dewhurst / Steller's jay (Cyanocitta stelleri) perched on branch. Grand Teton National Park, Wyoming, USA. September.

Oscar Dewhurst / Clark's nutcracker (Nucifraga columbiana) feeding on Pine (Pinus sp) seeds. Grand Teton National Park, Wyoming, USA. September.

Oscar Dewhurst / Killdeer (Charadrius vociferus) stretching wings. Yellowstone National Park, Wyoming, USA. September.

Ann & Steve Toon / Tawny eagle (Aquila rapax) in flight, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Warthog (Phacochoerus africanus) with Redbilled oxpecker (Buphagus erythrorhynchus), Zimanga game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Warthog (Phacochoerus africanus) drinking, with redbilled oxpeckers (Buphagus erythrorhynchus), Zimanga game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Warthog (Phacochoerus africanus) drinking, with redbilled oxpecker (Buphagus erythrorhynchus), Zimanga game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Greenbacked heron (Butorides striatus) peering from perch, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Woolly-necked stork (Ciconia episcopus) with frog, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Woolly-necked stork (Ciconia episcopus), Zimanga private game reserve, KwaZulu-Natal, South Africa.